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The University of the State New York 


New York State Museum 
sand ANNUAL REPORT 
1918 


VOLUME 2 


REPORT OF THE DIRECTOR 1918 
AND 
APPENDIX 2 


inns AMY 


ALBANY 
THE UNIVERSITY OF THE STATE OF NEW YORK 
1920 


THE UNIVERSITY OF THE STATE OF NEW YORK 


Regents of the University 
With years when terms expire 


(Revised to February 10, 1921) 


Piiny T. SExTON LL.B. LL.D. Chancellor -. - - Palmyra 
ALBERT VANDER VEER M.D. M.A. Ph.D. LL.D. 

Vice Chancellor Albany 
CHESTER 5S. Lorp M.A. LL.D. - -.- - - - - Brooklyn 
ADDEBERE WiloommcleaD =.=) 9 Vee = ee Se eB iittal@ 
CHARLES B. ALEXANDER M.A. LL.B. LL.D. Litt.D. Tuxedo 


WALTER Guest KeLitocGc B.A. LL.D. - - - - - Ogdensburg 
MANES WoVRNESBeAe lel. IDE = =o ner ta = = | New Mork 
HERBERT L. BripGMAN M.A. LL.D. - - - - - - Brooklyn 
THomas J. ManGAn M.A. - --- - - - - - - - Binghamton 
Warrant aN VICAR =) 5 SS Ee = = Yonkers 
Wittmam Bonpy M.A. LL.B. Ph.D: = - - =- = - New York 
WILLIAM P. BAKER B.L. - - - = - - - - - Syracuse 


Acting President of the University and Commisssioner of Education 


FRANK B. GILBERT B.A. LL.D. 


Assistant Commissioner and Director of Professional Education 


Aucustus S. Downine M.A. Ph.D. L.H.D. LL.D. 


Assistant Commissioner for Secondary Education 
CHARLES F. WHEELOCK B.S. LL.D. 
Assistant Commissioner for Elementary Education 


GEORGE M. WiLtEey M.A. Pd.D. LL.D. 


Director of State Library 


James I. WYER M.L.S. Pd.D. 
Director of Science and State Museum 


Joun M. Crarkre D:Se. LL.D. 


Chief and Directors of Divisions 
Administration, Hrram C, CasE 
Archives and History, JAMES SULLIVAN, M.A. Ph.D. 
Attendance, JAMES D. SULLIVAN 
Examinations and Inspections, AVERY W. SKINNER B.A. 
Law, FRANK B. GILBERT B.A. LL.D., Counsel 
Library Extension, WILLIAM R. Watson B.S. 
Library School, Epona M. SANDERSON, B.A. B.L.S. 
School Buildings and Grounds, FRANK H. Woop M.A. 
School Libraries, SHERMAN WILLIAMS Pd.D. 
Visual Instruction, ALFRED W. ABRAMS Ph.B. 
Vocational and Extension Education, Lewis A. WILSON 


yand ANNUAL REPORT 


OF THE 


NEW YORK STATE MUSEUM 


VOLUME 2 


To the Legislature of the State of New York 
We have the honor to submit herewith, pursuant to law, as the 72nd 
Annual Report of the New York State Museum, the report of the Director, 


with appendixes. 
Puiny T. SEXTON 


Chancellor of the University 

JouN H. FINLEY 
President of the University and 
Commissioner of Education 


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Appendix 2 
Museum Memoir 15 
Part 1 


Wild Flowers of New York 


The University of the State of New York 


> PA Weer Seneca 


JoHN M. CLARKE, Director 


Memoir 15 


WILD FLOWERS OF NEW YORK 


DUNT WO Paks 


BY 


HOMER D. HOUSE 


State Botanist 


Part © 


TABLE OF CONTENTS 


PAGE PAGE 
Letter of Communication............ 3 | Descriptions(Of Species». 4-4 35 
Introduction aa ia. joc eae 9) | Listrof lllustrations (pant i2) ee. -cee 325 
knots Swen sooovsocudccngdoowae TOs pebncl exal (pants) Meese rer ter ieee 341 
ALBANY 


THE UNIVERSITY OF THE STATE OF NEW YORK 
1918 


THE UNIVERSITY OF THE STATE OF NEW YORK 
Regents of the University 
With years when terms expire 
(Revised to July 1, 1918) 


1926 Priny T. SExToN LL.B. LL.D. Chancellor — -. — Palmyra 
1927 ALBERT VANDER VEER M.D. M.A. Ph.D. LL.D. 
Vice Chancellor Albany 


1922 CHESTER S. Lorp M.A. LL.D. - Brooklyn 
1930 WILLIAM NottTincHaAM M.A. Ph.D. LL.D. -— —- — Syracuse 
1921 FRANCIS M. CARPENTER = Mount Kisco 
1923 ABRAM I. Erxus LL.B. D.C.L. New York 
1924 ADELBERT Moor LL.D. - Buffalo 

1925 CHARLES B. ALEXANDER M.A. LL.B. LL.D. Litt.D. Tuxedo 

1919 JoHN Moore LL.D. = Elmira 

19628 WAULER GUEST Kritoce BA) LED. — — — — Ogdensburg 
1920 JAMES ByRNE B.A. LL.B. LL.D. = New York 
1929 HerBerRT L. BRIDGMAN M.A. Brooklyn 


President of the University and Commissioner of Education 
Joun H. Fintey M.A. LL.D. L.H.D. 
Deputy Commissioner and Assistant Commissioner for Elementary Education 
Tuomas E. FinEGAN M.A. Pd.D. LL.D. 
Assistant Commissioner and Director of Professional Education 
Aucustus S. Downinc M.A. L.H.D. LL.D. 
Assistant Commissioner for Secondary Education 
CHARLES F. WHEELOCK B.S. LL.D. 
Director of State Library 
James I. WYER, Jr, M.L.S. 
Director of Science and State Museum 
Joun M. CrarkeE D.Sc. LL.D. 
Chiefs and Directors of Divisions 
Administration, Hiram C. CASE 
Agricultural and Industrial Education, LEwis A. WILSON 
Archives and History, JAMES SULLIVAN M.A. Ph.D. 
Attendance, JAMES D. SULLIVAN 
Educational Extension, W1LL1IAM R. Watson B.S. 
Examinations and Inspections, GEORGE M. WiLEy M.A 
Law, FRANK B. GILBERT B.A., Counsel 
Library School, FRANK K. WALTER M.A. M.L.S. 
School Buildings and Grounds, Frank H. Woop M.A. 
School Libraries, SHERMAN WILLIAMS Ph.D. 
Visual Instruction, ALFRED W. ABRAMS Ph.B. 


LETTER OF COMMUNICATION 


New Vork State Museum 


January 30, 1918 
The Honorable John H. Finley 


President of the University 
SIR: 
The scientific survey of this State, established in 1836 under the title 


“ The Natural History of New York,’’ embraces in its monumental reports 
two volumes treating of the flora of the State. These volumes, prepared 
by the distinguished botanist, John Torrey, bear the inscription: Flora 
of the State of New York; Comprising Full Descriptions of All the Indigenous 
and Naturalized Plants Hitherto Discovered in the State, with Remarks on 
Their Economical and Medical Properties (1843). The species described 
in this work were entirely of the phenogamous or flowering plants. Until 
that time no summary of the New York flora had been brought together; 
and the service rendered to the people of the State by the publication of 
this compendium was of a high order and was received with enthusiastic 
appreciation. Doctor Torrey’s books served the needs of the time and 
expressed the state of its knowledge of the New York flora. 

Seventy-five years have passed, and in that long stretch of time botan- 
ical science has grown widely and apace. The field of cryptogamic botany, 
that which deals with the flowerless plants, the mushrooms, mosses, lichens 
and their kind, was not entered in these early reports; it was obscure and 
little understood; its mostly inconspicuous growths did not attract the 
eye or invite the observer; nor were its important relations to the economy 
of the community even suspected. 

The early official botanical investigations of the State were formally 
terminated by the publication of John Torrey’s reports. Not till 1867 did 
the need of continuous official attention to this department of science meet 

3 


4 NEW YORK STATE MUSEUM 


the recognition of the Regents of the University. In that year Charles 
Horton Peck was designated to take charge of such botanical collections 
as had accumulated in the State Museum, and not long thereafter Mr Peck 
was officially appointed the State Botanist. To the botanical service of 
the State Mr Peck thereafter dedicated himself without reserve for the 
rest of his long life. He added much to the store of knowledg» of the flow- 
ering plants, but the veiled world of the flowerless plants the more invited 
him and to it he specially gave his labors; leaving behind him a harvest of 
knowledge of them and a repute for his intricate researches which ranks 
him high on the roll of great botanists. Doctor Peck spared no effort, 
however, to increase the store of knowledge of all the flora of the State and 
he is the creator of the large state herbarium. After fifty years of unstinted 
devotion to his science and to his State, Doctor Peck fell asleep in honor, 
in the year 1917. 

Since the date of Torrey’s report, the flowering plants have been the 
subject of study in all parts of the Commonwealth. Botanical societies 
and local students have multiplied; records have grown; the demand for 
information has greatly increased; but there has been no reliable exposition 
of such information accessible to these students. 

It has been with this purpose of meeting a wide demand and of setting 
forth with such excellence as present knowledge and perfected modes of 
illustration could afford, that the present work, The Wild Flowers of New 
York, has been projected. The undertaking, bound to be an arduous one, 
has not been entered upon hastily. The advice of the leading botanists 
of this State and country was sought as to its timeliness, its scope, mode 
of presentation and illustration. The interested public will find it to be 
not a highly technical guide, couched in closely analytical descriptions, but 
a comparatively brief text, untechnical so far as the theme permits, accom- 
panied by color illustrations made from the growing plants. The present 
State Botanist, Dr Homer D. House, is the responsible author of the work; 
he has not only prepared the text and its arrangement, but has supervised 
in detail the color photography; he has accompanied the photographers 


WILD FLOWERS OF NEW YORK 5 


into the field on every visit and has selected every subject which is here 
reproduced. 

The color illustrations must speak for themselves. They have been 
executed with most painstaking care by the quadricolor process, and the 
living and growing plants have been reproduced in their colors as near to 
those of nature as now seems possible. Credit for the printing and binding 
of the work should be given to the J. B. Lyon Company of Albany, but 
the Matthews-Northrup Company of Buffalo and the Zeese-Wilkinson 
Company of New York are to be accredited with the quality and making 
of the color plates. 

As such, then, these volumes are given to the people of the State and 
as such, we believe, they would have this service rendered. 

Very respectfully 
JOHN M. CLARKE 


Director 


One who is upon the gray ocean at this season of the year when, in 
the woods and at the roadsides in the State of New York, the wild flowers 
are beginning to redeem their promises of life, appreciates as never before 
how much these quiet, persistent pioneers of the fields contribute in scent, 
color and form to the making of that which is summed up in the name 
New York; even as the heather to the making of that country whose head- 
lands are now dimly emerging from the level sea. The sight of a spray of 
these native flowers, such as many a page in this book carries, would be 
as a twig borne back in ancient times to the ark —a sign that, though the 
flood of war has overwhelmed many valleys, the elemental processes of life 
go forward undisturbed in the “ Empire State.” Whatever the economic 
value or imputed harm of these aborigines, first settlers, later immigrants 
and vagrants which together constitute the Flora of the State, it is desirable 
from every point of view, since they are our near, most welcome but some- 
times intrusive neighbors, that we should know their faces, their habits and 
their capacities for good or warning of ill. Itisa great realm of life within 
the State of which the State as a whole should acknowledge the possession. 

I have unusual satisfaction in finding it my official opportunity to say 
an introductory word to this notable and distinguished work, because it .is 
the record of a possession which the Director of the State Museum, Dr John 
M. Clarke, has enabled the State to make. It has a great practical value, 
but it has another value in making perennial and keeping in perpetual 
domestic bloom, in home and schoolhouse and library, flowers that blossom 
but a few days or weeks in the wild state in which they have been so skil- 
fully and sympathetically discovered by Doctor House. I am proud that 
the State has made possible such a publication and that The University 
of the State of New York has been able to execute the commission with 
such success. 


. 
———————— 


On the Atlantic Ocean President of the University 


May 19018 7 


; e121 
a con iS 
“any 


Fe 


Arwen: 


WILD FLOWERS OF NEW YORK 


INTRODUCTION 


The State of New York possesses a large variety of herbaceous and 
shrubby plants with conspicuous flowers, which may be classed under the 
rather broad and indefinite term of “‘ wild flowers.’’ For the purposes of 
this work only a few of the shrubs, such as the Mountain Laurel, Azalea 
and Labrador Tea have been included, the thought being to present mainly 
herbaceous plants with conspicuous flowers. 

Anyone who has observed the natural vegetation in such unlike parts 
of the State as the salt marshes and pine barrens of Long Island, the higher 
Adirondack and Catskill mountains or the woodlands of the western counties 
must have been impressed by the obvious difference in the wild flowers of 
those several sections, and especially by the fact that very few of the 
wild flowers which bloom between early spring and late autumn in the 
Adirondacks are to be found on Long Island. 

Such differences in the character of the vegetation of widely separated 
portions of the State are explained partly by soil conditions and partly by 
differences in climate. Located with the ocean at one side and the great 
inland lakes at the other, the State is favored by conditions of atmospheric 
moisture (relative humidity, rainfall and snowfall) which make it climati- 
cally a forest region, and hence favorable for a luxuriant variety of herbace- 
ous and shrubby plants; a region in which forests would naturally dominate 
all other vegetation if not cut down. The temperature conditions along 
the southern coast of the State are modified by the ocean, and to some 
extent on the west by the Great Lakes, while the elevated mountain masses 
of the Adirondack and Catskill regions produce cooler summers and shorter 
growing Seasons. 

I am indebted to Mr Edward A. Eames of Buffalo for photographs and 
autochromes of certain orchids, to Mr G. A. Bailey of Geneseo, and Mr O. O. 
Nylander of Caribou, Me., for additional photographs and to Mr Louis R. 


Robbins, former assistant to State Botanist, for assistance in the preparation 
of the text and illustrations for the chapter on Plant Structure. 


9 


10 NEW YORK STATE MUSEUM 


PLANT STRUCTURE! 


No one who loves plants, either cultivated or wild, has failed to note 
how they differ from one another in shape, size, color and arrangement of 
the flowers, the leaves and other parts. These features are essentially the 
same in all individuals of a given species, but differ greatly in individual 
plants not belonging to the same species or variety. 

To express these differences requires a terminology that is familiar 
largely only to those who have studied botany. The fact that it is practically 
impossible to describe a plant accurately without the use of a certain number 
of these special terms is a great impediment to a broad familiarity with 
our wild flowers on the part of all those who would like to study them with 
the aid of botanical guides. Because of the great variety of flowering 
plants and the minuteness of the flowers on many of them, the difficulty of 
identification is even greater than that connected with the study of bird 
and animal life. 

The following brief summary of the terms necessary to an accurate 
description of a flowering plant, taken alone, means little to the average 
reader. Taken in connection with the plants as they are found growing 
and carefully studied, these terms, as soon as understood, place one on a 
footing of easy familiarity with the wild flowers, so that an accurate descrip- 
tion as given in books means something definite and enables one to decide 
if the given description applies to the plant under consideration. Supple- 
mented by illustrations, a study of the terminology used enables the student 
to acquire a much wider knowledge of our wild flowers. 


Leaves 
The leaf is an essential organ of all plants which live independently, 
that is, are not parasitic upon other plants (like the Dodder) or saprophytic 
upon dead plant remains (like the Indian Pipe). The leaf manufactures 


‘The cuts in this section are adapted from Gray’s Lessons in Botany. Copyright by 


Asa Gray. Reproduced by permission of the American Book Company, publishers. 


WILD FLOWERS OF NEW YORK II 


food for the plant, gives off excess water (transpiration) and is the breathing 
organ of the plant. To accomplish these functions the leaf is built up by 
a complex arrangement of cells and is variously modified in different groups 
of plants to meet the external conditions of environment and competition 
by other plants. 

The parts of a leaf are designated as 
blade, petiole and stipules. The leaf blade 
(figure 1A) is the broadly expanded portion, 
although in some species the leaf blade is 
very narrow or even threadlike. The petiole 


(figure 1B) is the stalk which supports the 
blade, and may be lacking in some cases, when 
the leaf is said to be sessile. The stipules (figure 1C) are small, leaflike 
organs at the base of the petiole, and are best typified by the rose leaf. 
Frequently the stipules encircle the stem at the base of the petiole and often 
they are entirely lacking or fall away so soon after the leaves expand that 
they are not found when the plant is in bloom. 

Terms of leaf outline: The various shapes of leaf blades may be 
expressed by the following terms: 

Subulate; awl-shaped, without visible expansion of blade, and usually 
tapering to the apex (figure 3). 


3 4 6 ul 
Linear, or ribbon-shaped; elongated and several times longer than 
wide (figure 4). 


2 NEW YORK STATE MUSEUM 


Lanceolate; in which the leaf blade is three times as long as wide, or 
longer, and broadest at or below the middle (figure 5). 

Oblong; in which the blade is somewhat longer than wide, broadest 
in the middle or with sides almost parallel (figure 6). 

Ovate ; shaped like an egg; that is, broadest below the middle or near 
the base (figure 7). 

Elliptical; rounded at both ends, somewhat longer than wide (figure 8). 

Orbicular or rotund; in which the blade is nearly or quite circular in 
outline (figure 9). 

Reniform; in which the blade is 
broader than long, with a heart-shaped 
base (figure 10). 

Deltoid; triangle-shaped, similar 
to ovate but conspicuously broadened 


at the base and pointed at the apex 


(figure I1). 


Consideration of a few leaf blades 
shows immediately that these terms 
are not always sufficient to express 
accurately the shape and we may have 
recourse to combinations of terms, such 

. as oblong-lanceolate, ovate-lanceolate 
10 11 (figure 13), etc. 
The shape of leaf blades which are 
broadest above the middle may be expressed by the following terms: 
Obovate; ovate in shape, but broadest near the apex or above the 
middle (figure 14). 
Oblanceolate; lanceolate in shape but broadest above the middle or 
near the apex (figure 15). 
Spatulate; in which the blade is oblanceolate or obovate in shape with 


the base conspicuously elongated (figure 12). 


WILD FLOWERS OF NEW YORK 13 


Terms applied to the apex of the leaf: 
Obcordate; broad and heart-shaped at the apex (figure 16). 
Emarginate; with a slight depression at the somewhat narrowed apex 
(figure 17). 
Retuse; terminating in a semicircular end, the center of which is 


somewhat indented (figure 18). 
Truncate; with a flat or abrupt apex (figure 19). 


12 13 14 15 


Acuminate; when the apex of the blade is longer than broad (figure 20). 
Acute; when the apex of the blade is about as broad as long (figure 21). 
Obtuse or blunt; when the apex is much broader than long (figure 22). 


14 NEW YORK STATE MUSEUM 
Mucronate; when the apex is terminated by a short blunt tip (figure 


22) 


Cuspidate ; when the tip of the blade is hard and stiff (figure 24). 


Terms applied to the base of the leaf: 
The terms truncate, acuminate, acute, obtuse (defined above) may 
also be applied to the shape of the base of the leaf blade, in addition to the 


following: 


\\26 


2 
} | 31 


30 


Cordate; heart-shaped (figure 25). 


32 


Cuneate, or wedge-shaped; when the sides of the leaf blade taper to 
an acute angle at the base (figure 27). 

Auriculate, when the depression at the base of the blade is deep and 
produces on either side conspicuous basal lobes (figures 28 and 32). 

Sagittate; when the basal lobes point downward like the head of an 
arrow (figure 30). 


Hastate; when the basal lobes are turned outward (figure 29). 


WILD FLOWERS OF NEW YORK I 


on 


Peltate; a rounded leaf blade with the petiole attached at or near the 
middle of the lower surface (figure 31). 

Perfoliate; when the bases of leaf blades meet and join around the stem 
of the plant (figure 26). 


Terms applied to the marginal segmentation of leaf blades: 


Sinuate; when the marginal lobes of the leaf blade present a wavy 
outline (figure 33). 


36 


Pinnately lobed; when the tissue between the veinlets is cut out nearly 
to the midrib of the leaf and the divisions are arranged like the pinnae of 
a feather (figure 34). 

Palmate; when the blade is deeply divided nearly or quite to a common 
base (figure 38). 


16 NEW YORK STATE MUSEUM 


Serrate (figure 35); when the margin is sharply toothed with coarse 
teeth, like a saw. When the teeth are rounded inward or are convex, the 
margin is said to be dentate (figure 37). When the 
margin is formed of rounded teeth it is said to be 
crenate (figure 36). If the teeth are very small, the 
diminutives of the above terms are used, namely, 
serrulate (figure 39), denticulate (figure 40) and 
crenulate (figure 41). 


Compound leaves: 

39 40 41 When a leaf possesses several divisions or seg- 
ments upon a common petiole or rachis, 1t 1s said to be compound. The 
distinction between a simple leaf, which is deeply divided, and a compound 
leaf, rests upon the presence of distinct articulation between the leaf seg- 
ment of the compound leaf and the petiole. Compound leaves may be 
pinnate (figure 42), when the leaflets are arranged on either side of a com- 
mon petiole (Ash, Rose, Walnut etc.) or palmately compound (figure 43), 
when the leaflets all join the petiole at its summit (Horse-chestnut). 


Leaf arrangement: 


Alternate, when the leaves are arranged one at a node and each leaf 
is opposite and above the preceding leaf; spirally arranged, when the nodes 


WILD FLOWERS OF NEW YORK 7, 


are not opposite; and opposite when the leaves are in pairs opposite each 
other on the same node. When several leaves are inserted on the same 
node they are said to be whorled or verticillate. 


Flowers 


The flower of a plant is a group of organs (figure 44) which exist for 
the purpose of producing seed. The parts of a flower fall into two general 
groups: those which ac tually function 
as seed producers (essential organs), 
and those which act as _ protective 
organs or organs for the attraction of 
insects (floral envelops or perianth). 
These might also be designated as non- 
essential organs, since they are lack- 
ing in certain flowers. 

The essential organs consist of 
two parts, the pistils (figure 44A) and 
the stamens (figure 44D, E), often 


designated as the pistillate and stami- 
nate parts of a flower; and when a 
flower contains only pistils it is called a pistillate flower, and when it 
contains only stamens it is called a staminate flower. 

When both stamens and pistils are present in the same flower, it is 
said to be perfect. If, in addition, the flower possesses the floral envelops, 
calyx and corolla, it is called a complete flower. Hence a flower which 
lacks any of these sets of organs is incomplete (that is, if it lacks either 
calyx or corolla); if it lacks either stamens or pistils it is imperfect. 


The perianth or floral envelopes: 


The ideal flower contains two sets of floral envelops, the calyx 
(figure 44F) and the corolla. In some flowers the corolla is entirely or 
partly divided into a certain number of divisions, each of which is called 


18 NEW YORK STATE MUSEUM 


a petal (figure 44G). They are usually but not always brightly colored. 
Subtending or beneath the corolla is the calyx, which is usually, but not 
always, green, and is likewise in many plants divided into a number of 
distinct parts or sepals. When an incomplete flower has but one set of 
floral envelopes, it is usually the petals (or corolla) which are lacking, and 
in such cases the calyx may be brightly colored and function as a corolla 


(a petaliferous calyx). 


The essential organs: 

The number and arrangement of stamens varies in different kinds of 
plants, but nearly always a stamen consists of a filament or stalk (figure 
44E), which bears at its apex the anther (figure 44D), or pollen-bearing 
sac. The shape of the anther, and the manner by which it dehisces, or 
opens to emit the pollen, likewise varies in different groups of plants. 

The pistil (figure 44A—C), or seed-bearing organ, consists of an ovary 
(figure 44A), stigma (figure 44C) and style (figure 44B). The ovary is at 
the base of the pistil and contains the 
ovules or eggs, which after fertilization 
ripen into seeds. The ovary usually con- 
tains several or many ovules, but may 
contain as few as a single ovule. The 
stigma is that part of the pistil which 
acts as a receptive organ for pollen in the 
process of pollination. Its surface is 
usually moist and minutely granular and 
its position and shape are dependent upon 


45 


made use of by the particular plant. The style connects the stigma and 


the mode of pollination (insects or wind) 


ovary. It may be long or short, slender or stout, or sometimes entirely 
lacking when the stigma is situated directly upon the ovary. 

The ovary itself may contain one or several chambers or cells (figures 
45-47), and very frequently the number of chambers in the ovary and the 


WILD FLOWERS OF NEW YORK 19 


lobes or divisions of the stigma bear a direct relationship to the number 
of petals, sepals and stamens. The term carpel (or carpophyllum) is used 
to designate the seed-bearing leaf. A carpel may be a pistil of itself, or it 
may be a constituent of a more complex pistil. In either case, a carpel 
is the homologue of a leaf. The surface within the ovary to which the 
ovules are attached is called the placenta. 

Simple pistils may be solitary, or several together on a common recep- 
tacle within the flower, as in the Buttercup. A compound pistil consists 
of two, three or more carpels united into one body. 

The apex of the flowering stem, which supports the flower, is 
designated as the receptacle. 


48 49 50 51 


Arrangement of flowers: 

Flowers are either solitary or clustered, but their arrangement varies 
in different kinds of plants, and may even vary to some extent in the same 
species. The arrangement or disposition of the flowers may be designated 
as the inflorescence. The following are the most frequent arrangements 
of flowers: 

Spike (figure 48), in which the flowers are arranged along the flowering 


20 NEW YORK STATE MUSEUM 


stem, and sessile (that is, without stalks) or with very short stalks (pedicels). 
When the flowering stem is naked (devoid of leaves) and rises directly 
from the root or crown of the plant, it is called a scape (figure 51). 

Raceme (figure 49), in which the flowers are arranged along a flowering 
stem and each flower possesses a distinct stalk or pedicel. The lower 
pedicels may be somewhat longer than the upper ones. 

Umbel (figure 52), when the flowers arise from the same point, which 
is usually the apex of the flowering stem or of a lateral flowering stalk, and 


54 55 


radiate like the rays of an umbrella. If the radiating stalks of such an 
inflorescence bear smaller umbels at their tips, it is called a compound 
umbel. 

Corymb |corymbose| (figure 54), when the branches of an inflorescence 
are of unequal length, but the lower or outer ones are longest so that they 
all form a flat-topped, or nearly flat-topped, cluster. 

Cyme (figure 55), when the flowers each terminate an axis or stem 


arising successively from a new axis or stem. 


WILD FLOWERS OF NEW YORK 21 


A spadix is a spikelike inflorescence with a fleshy stalk and with sessile 
flowers; the floral leaf or bract which subtends it or surrounds it partially 
is called the spathe (Skunk Cabbage, Wild Calla). 

A panicle, or compound raceme (figure 50), is formed by the arrange- 
ment of flowers along the plant stem, similar to a raceme, but each flower 
stem has two or more branches. 

Head (figure 53), an arrangement of flowers compactly on a common 
receptacle and surrounded by bracts (involucral bracts). 


Modification and arrangement of the perianth: 


Among the simpler groups of flowering plants the perianth is wanting, 
as in the Cat-tail and Willow. In the Sweet Flag, Oak and others, the 
perianth consists of a few scales, but in the higher plants, the perianth 
appears as a conspicuous portion of the flower, as in the Lily. Finally, as 
in the Rose family, there appears a clearly differentiated calyx and corolla. 

In the simpler types of flowers, the sepals, petals and the stamens 
arise at the top of the receptacle. Such flowers are called hypogynous, 
meaning the insertion of these parts below the ovary (figure 56). 

When the basal portion of the receptacle is continued upward, forms 
a cup-shaped growth around the ovary and bears the sepals, petals and 
stamens upon its margin, the flower 1s called perigynous (figures 57 and 59), 
meaning the insertion of the parts of the flower around the ovary. 

Frequently the growth of the receptacle adheres to the ovary, and the 
sepals, petals and stamens appear to arise from above the position of the 
ovary, in which case the flower is called epigynous (figure 58), meaning 
above the ovary. 

There may be varying degrees of cohesion or union of the parts of one 
or both of the floral envelopes (perianth). When the sepals are united with 
each other the calyx is said to be gamosepalous, while a gamopetalous 
corolla (figures 62, 63, 64 and 65) refers to a union of the petals, as in the 
flower of the Morning-glory. 

The degree of coalescence or union of parts of a gamopetalous corolla 


NEW YORK STATE MUSEUM 


WILD FLOWERS OF NEW YORK 23 


varies in different flowers. When the calyx or corolla is divided almost to 
the base it is said to be parted (figure 63); when divided to about the middle 
it is said to be cleft (figure 64); when still less separated it may be said to 
be lobed or toothed (figure 65); or if entire on the margin it is said to be 
entire (figure 62). 

When the parts of each set of organs of a flower are alike or equal in 
size, the flower is said to be regular, which means that the petals are alike, 
the sepals are alike and the stamens are alike. A symmetrical flower is 
one in which the sepals, petals and stamens are of the same number; 
unsymmetrical when there are unequal numbers in each cycle, that is, 
an unequal number of sepals, petals or stamens. 

Certain groups of plants may often be recognized by the form of the 
corolla of some of its members. This character seems to be quite constant 
and the names of several large or important families of flowering plants 
are derived from this source. Of these groups we may mention the 
cruciferous (figure 60) type of flower of the Mustard family (Cruciferae), 
in which there are four spreading petals forming a cross, as in the flower 
of the Spring Cress (Cardamine bulbosa); the labiate corolla 
(figure 66) of the Mint family (Labiatae) in which the corolla is more or 
less two-lipped; the papilionaceous type of flower (figure 61) of the Pea 
family (Leguminosae), in which the petals are characteristically grouped 
into two lateral (wing) petals, a single upper (banner) petal and a pair of 
lower petals, often more or less united to form the keel. 


Stamens: 


The general characteristics of the stamens have already been described. 
In the stamens, as in the case of the petals and sepals, the number and 
arrangement are subject to great variation in different kinds of plants. 
Monandrous refers to a flower with a single or solitary stamen; polyandrous 
to a flower containing several stamens. The stamens may be monodelphous, 
in which the filaments are united into a tube, as in the Wild Lupine (figure 
67), or the stamens may be diadelphous (figure 68), which means two sets 


NEW YORK STATE MUSEUM 


WILD FLOWERS OF NEW YORK 


25 
of united stamens. In this form of arrangement there may be a union of 
the filaments of all the stamens except one, which is a common diadelphous 
arrangement of stamens in many of the species of the Bean family. When 
there are several sets of united stamens, the arrangement is said to be 
polydelphous. 

Adnation or union of the stamens with other parts of the flower is of 
frequent occurrence, and the terms employed depend upon the degree of 
adnation, or the absence of it, namely, hypogynous (meaning beneath the 
pistil), applied to parts, including stamens, which are inserted or borne 
on the receptacle of the flower (figure 56). This is the absence of adnation 
and indicates an unmodified type. Perigynous (around the pistil) implies 
an adnation which carries up the apparent origin or place of insertion of 
the parts of the flower to some distance above or away from the receptacle 
and thus placing the insertion around instead of beneath the pistil (figure 57). 
Epigynous (on the pistil), where the adnation is complete to the very top 
of the ovary (figure 58). 

When the stamens are borne upon the corolla, or upon the tube of 
the corolla, they are said to be epipetalous (figure 69), and when they are 
borne upon the pistil, as in the Orchid family, they are said to be gynandrous. 

The most important part of a stamen is the anther (figure 44D), which 
contains the pollen. It normally consists of two lobes or sacs; but as each 
sac is often, and in most of our common flowers, divided into two cavities, 
it appears to possess in such instances four pollen sacs. For the discharge 
of the pollen, the cells of a normal anther open along a definite line, usually 
extending from top to bottom. This suture or line of dehiscence may be 
lateral or marginal, or centrally located. 

In the genus Solanum, to which the Potato belongs, in most members 
of the Heath family (Ericaceae), in Polygala, and certain other species, the 
anther cells open only by a hole or pore (figure 71). In the Blueberry, 
Cranberry etc. the pore-bearing tip of the anther cell is prolonged con- 
siderably, often into a slender tube. In the Barberry, and in most other 
members of that family, and in the Lauraceae, the whole face of each anther 


26 NEW YORK STATE MUSEUM 


cell separates by a continuous line, forming a kind of door, which is attached 
at the top, and turns back, as if on a hinge; and the anther is said to open 
by uplifted valves (figure 72). In the Sassafras and certain other members 
of the Lauraceae, each lobe of the anther opens by two smaller valves, like 
trapdoors. 

The attachment of the anther to the filament (or stalk) presents three 
different modes, frequently connected 
by gradation: Innate (figure 70), in 
which the anther is a direct continua- 
tion of the axis of the filament, the 
cells usually opening by marginal slits, 
and the lobes or cells of the anther 
project neither inward nor outward; 
adnate (figure 73),in which the anther 
is a direct continuation of the filament 
but having the anther cells adherent 
to the anterior or posterior face of the 
filament; the Wild Ginger (Asarum) 
furnishes a good example of this, on 


account of a prominent prolongation 
5 of the connective or tip of the fila- 

ment (figure 74); versatile (figure 75), 
when the anther is attached at some part only of its back or front to the 
tip of the filament, on which it lightly swings when the pollen is discharged ; 
examples of this are seen in members of the Lily family, the grasses, Evening 


Primrose (Oenothera biennis) and others. 


Pollination 
The structure of most flowers affords an excellent indication of the 
device used for the transference of pollen from one flower to another 
(pollination), Long ago it was assumed that Nature wished no flower to be 
fertilized by its own pollen, but in the light of present knowledge we know 


WILD FLOWERS OF NEW YORK 27 


this is not wholly true. The subject of pollination of flowers by insects 
received a great light through the investigations of Charles Darwin and the 
publication in 1862 of his well-known book on the fertilization of orchids 
by the aid of insects. 

As we understand the matter today, it appears that flowers are habitu- 
ally intercrossed (flowers of the same species), and that there are manifold 
structural adaptations which secure or favor this interchange of pollen. 
Separation of sexes (stamens and pistils) is a direct adaptation to cross- 
pollination, rendering it necessary between individuals with dioecious 
flowers, and favoring it in most plants with monoecious and polygamous 
flowers. Strictly, close fertilization can take place in hermaphroditic 
flowers only. 

Flowers depend upon certain external agencies for the transference 
of pollen from one flower to the flower on another plant. These agencies 
are wind (anemophilous flowers) and insects (entomophilous flowers). 
Other agencies are of minor importance, although water must be considered 
in connection with some aquatic plants. 

Wind-pollinated flowers are mostly dull in color, destitute of odor and 
nectar, since these qualities attract insects. Wind-pollinated flowers 
usually have the sexes separated, the flowers borne in great abundance 
and have very light pollen. Most of our common trees (the Pines, Oaks, 
Hickories etc.) depend upon wind for the transference of pollen, as do also 
the grasses, sedges, Plantain and others. 

Insect-pollinated flowers are correlated with showy coloration (includ- 
ing white, which is most showy at dusk), odor or secretion of nectar. 
Structural adaptations of the flower in reference to insect visitation are 
wonderfully various, and most of these are found upon investigation to 
favor, or often to necessitate, cross-pollination. The range of these varia- 
tions is too extensive to be treated here. Literature upon this subject is 
easily available and most textbooks of botany contain chapters upon the 
subject. 

After pollination the pollen grain germinates upon the surface of the 


28 NEW YORK STATE MUSEUM 


stigma, sends a tube down through the tissue of the stigma and style and 
discharges into the ovule a male nucleus which unites with a nucleus in the 
embryo sac of the ovule, fertilizing the ovule, and stimulating its develop- 
ment into an embryonic plant. By a process of hardening of the coats of 
the ovule its development is arrested and the seed is produced. 


The Fruit 

The fruit of a plant (in the case of our flowering plants) consists of the 
matured pistil (or gynoecium), including also whatever parts of the perianth 
or other floral organs may be joined to it. Fruits are of various degrees 
of simplicity or complexity, and may consist of a matured simple ovary, 
a cluster of such ovaries, at least when they are somewhat coherent, or 
a ripened ovary with calyx and other floral parts consolidated with it. 

The pericarp, or seed vessel, is the ripened ovary and should therefore 
accord in structure with the ovary from which it is derived. In the 
development of a simple ovary into a simple fruit certain alterations some- 
times take place, either by the abortion or obliteration of certain parts, or 
by accessory growth. The dehiscence is the method by which a pericarp 
opens to discharge its seeds and may be regular (normal) or irregular 
(abnormal). The word “ pod ”’ is frequently applied to dehiscent pericarps. 

A capsule is a dehiscent pericarp formed of two or more carpels. Such 
carpels are septicidal (figure 80) when the dehiscence is such that the carpel 
is divided into its constituent carpels. Members of the St John’s-wort 
family afford a good example of this method as do also Rhododendron and 
Kalmia. Carpels are called loculicidal (figure 79) when each of the compo- 
nent carpels splits down its dorsal suture, as in Iris, Hibiscus, Oenothera etc. 

Kinds of fruits. For ordinary purposes it is sufficient to classify fruits 
into four classes: 

[ Simple fruits, those which result from the ripening of a single pistil. 

2 Aggregate, those of a cluster of carpels of one flower crowded into 


a mass. 


WILD FLOWERS OF NEW YORK 29 


3 Accessory fruits, where the principal mass consists of the surround- 
ings or support of either a simple or an aggregate fruit. 

4 Multiple or collective fruits, formed by the union or compact 
aggregation of the pistils of several flowers. 


I Simple Fruits 

Upon the basis of texture, simple fruits may be designated as dry 
fruits, stone fruits and baccate fruits. 

Dry fruits which are dehiscent: 

Follicle (figure 78), a pod formed by a simple pistil, and dehiscent 
along one line (suture, and almost always the inner or ventral suture), 
as in the Columbine, Marsh Marigold, Milkweed and Dogbane. 

Legume (figure 77), a pod formed 
of a simple pistil which is dehiscent 
by both sutures, so dividing it into 
two pieces or valves. The fruits of 
the Bean or Pea family are of this 


a" Wee 


myn 


sort. Some members of this family 


(Meibomia), however, have legumes 
reduced to indehiscent achenes, joined 


together end to end, and to which a 
special term “loment” (figure 76) is 


78 


applied. 
Capsule (figures 79 and 80), a pod 
or dehiscent fruit, of any compound pistil. The modes of regular dehiscence 


are mentioned above in the paragraph on dehiscence, and it remains here 
to describe two modifications of the capsule, namely, the pyxis, in which 
the dehiscence is along a circular line, cutting off the upper part as a lid, 
examples of which are seen in the common Plantain, Purslane and Hen- 
bane, small plants or weeds not illustrated in this work; and the silique, 
a narrow, two-valved capsule, with two parietal placentae, from which the 


30 NEW YORK STATE MUSEUM 


valves separate in dehiscence, as in the Mustard family, where there is 
usually a false partition stretched across between the two placentae. 

Dry fruits which are indehiscent: 

Samara, an indehiscent, one-seeded fruit 
provided with a wing. In the Ash, the wing 
is terminal; in the Elm, the wing surrounds the 
body of the pericarp; and the Maple fruit is a 
double samara or pair of such fruits. 

Achene (figures 81-88), a general term for 
all one-seeded, dry and hard, seedlike fruits. 
The best examples are the fruits of the Butter- 
cup, Anemone, Clematis and Avens. The style 
sometimes remains on the fruit as a long and 
feathery tail (Dandelion, figure 85), and in others 
merely as a short hook (Buttercup, figures 86 
and 87). Inthe Compositae (Sunflower family) 
the tube of the calyx is joined with the surface 
of the ovary, and its border or upper edge 


appears as a crown or cup, or a set of teeth or of 
scales, or very often as a tuft of bristles or hairs, called the pappus (figures 
82-84, 88). 

Utricle, a dry achenelike fruit with a thin and bladdery loose pericarp, 
like that of the Goosefoot (Chenopodium). 

Caryopsis or grain, differs from the achene in having the seed completely 
filling the cell and its thin coats firmly consolidated throughout with the 
very thin pericarp. This term is applied to the fruits of the grass family, 
including Indian corn and all other cereals. 

Nut, a hard one-celled and one-seeded, indehiscent fruit which finds 
its best examples in the fruit of the Hazel, Beech, Oak, Chestnut etc. The 
smaller nutlike fruits of the Borrage family and of the Mint family are 


usually called nutlets. 


WILD FLOWERS OF NEW YORK 31 


Stone fruits: 

Drupe (figures 90 and 92), of which the best examples are the fruit of 
the Cherry, Plum, Peach etc., are one-seeded or rarely two-seeded, in the 
ripening of which the outer portion of the pericarp becomes fleshy or pulpy 
and the inner portion becomes much hardened. The term is also commonly 
applied to similar fruits of the Hackberry, Cornus, Rhamnus etc. In the 
case of the Blackberry (figure 89) and Raspberry, the several pericarps of 
the aggregate fruit are called drupelets. 


Pome (figure 91), the name of the fruit of the Apple, Pear, Quince etc., 
which are fleshy fruits, composed of two to several carpels, of parchment- 
like texture (or hard in the Thorn Apples), inclosed in flesh which has 
developed from the inclosing calyx and receptacle. Indeed, the fruit of 


‘ 


the Thorn Apple might well be called a “ several-seeded drupe.” 
Pepo, or Gourd-fruit, a type of fruit typified by the Melon, Squash, 


Cucumber, Gourd and other members of that family. 


NEW YORK STATE MUSEUM 


Ww 
i) 


Berry [baccate] (figure 93), a simple fruit in which the pericarp is fleshy 
throughout and without a hardened inner coat. The fruit of the Grape, 
Currant, Gooseberry, Cranberry, Banana and Tomato furnish good 


examples. 


2 Aggregate Fruits 


Aggregate fruits are those in which a cluster of carpels, all belonging to 
one flower, are crowded on the receptacle into one mass, as in the Black- 
berry (figure 89) taken as a whole. They may be aggregates of any kind 
of simple fruits. But when dry and not coherent, the mass would simply 
be described as a head or spike of carpels (or achenes, as in Buttercup, 


Anemone etc.). 


WILD FLOWERS OF NEW YORK 33 


3 Accessory Fruits 

Accessory fruits are those in which some conspicuous part of the fruit 
is derived from some portion not organically connected with the ovary or 
pistil. This part might be called a pseudocarp, and this condition may 
occur either in simple, in aggregate, or in multiple fruits. The Winter- 
green (Gaultheria procumbens) affords a good example (figures 94 
and 95), the fleshy part of the fruit being the enlarged calyx. Likewise 
the torus, although not conspicuous, may be said to be an accessory part 
of the fruit of the Blackberry, being the fleshy or pulpy center of the 
fruit. In the Strawberry it is very conspicuous and comprises the sole 
edible part of the fruit, the achenes or true fruits being dispersed over the 
surface and comparatively insignificant. 


4 Multiple or Collective Fruits 
Multiple or collective fruits are those which result from the aggregation 
of several flowers into one mass. The simplest of these is the fruit of the 
Partridge Berry (Mitchella repens) and certain Honeysuckles (Loni- 
cera) formed of the ovaries of two blossoms united into one fleshy fruit. 
More typical examples of this are seen in the Pineapple fruit, the Mulberry 
and others. 


WILD FLOWERS OF NEW YORK 
DESCRIPTIONS OF SPECIES 
Cat-tail Family 


Typhaceae 
Broad-leaved Cat-tail 
Typha latifolia Linnaeus 
Figure I 

A marsh or aquatic plant, usually growing in thick colonies from 
creeping perennial rootstocks provided with fibrous roots. Stems stout, 
round in cross-section, glabrous, 4 to 8 feet high. Leaves numerous, 
linear, flat, swordlike, sheathing the stem at the base and rather stiffly 
ascending. Flowers monoecious, that is, staminate and pistillate flowers 
separate but on the same plant; densely crowded into terminal spikes; 
the staminate spikes uppermost and scarcely or but slightly separated 
from the dark brown or nearly black pistillate spike, each 3 to 12 inches 
long and often an inch or more thick. Perianth of the individual flowers 
composed merely of bristles which subtend two to seven stamens (in the 
staminate spike), or a small, short-stalked ovary (in the pistillate spikes). 
Mingled among the stamens and pistils are bristly hairs, and among the 
pistillate flowers many sterile flowers with clavate tips. The fruit consists 
of many small nutlets, surmounted by the persistent bristles which aid in 
wind dissemination of the seeds when the head of fruit breaks up. 

Common everywhere in marshes and swamps, and also in Europe and 
Asia. Flowering in June and July; fruit ripe in August and September, 


frequently persistent until the following spring. 


The Narrow-leaved Cat-tail (Typha angustifolia Linnaeus), 
is more abundant in marshes along the coast, but is sometimes found inland. 
The leaves are narrower than those of the preceding species, being one- 
sixth to one-half of an inch wide; spikes lighter brown in color, not so 
thick, and the staminate and pistillate spikes separated from one another. 


35 


36 NEW YORK STATE MUSEUM 


Figure I 


Broad-leaved Cat-tail 


(Typha latifolia Linnaeus) 


WILD FLOWERS OF NEW YORK 37 


Water Plantain Family 
Alismaceae 
Broad-leaved Arrowhead 
Sagittaria latifolia Willdenow 
Plate 1 

A perennial aquatic herb with thickened base and numerous long, 
fibrous roots. Leaves long petioled and extremely variable in form and 
size, sometimes wider than long and obtuse, sometimes linear-lanceolate 
and acuminate at the apex; the basal lobes of the leaf blades one-quarter 
to one-half as long as the blade. Flowers monoecious or sometimes 
dioecious, pediceled and borne near the summits of the scapes in verticels 
of three, the staminate usually uppermost, each verticel subtended by 
three bracts. Calyx of three persistent sepals. Stamens numerous; 
anthers two-celled, dehiscent by lateral slits. Pistillate flowers with 
numerous distinct ovaries and sometimes with imperfect stamens. The 
ovaries ripen into a globose or compact head of achenes, each achene 
broadly winged on both margins, with a beak about one-third its length 
and horizontal or nearly so. 

Common in shallow water almost everywhere and offered by dealers 
in native plants for colonizing lily ponds and shallow waters. Such situa- 
tions are scarcely complete without its presence. Flowering from July to 
September. 

There are several closely related species such as Sagittaria cuneata 
Sheldon, with a minute beak to the achene, which is erect over the ventral 
wing; Sagittaria pubescens Muhlenberg, which is strongly pubescent; 
Sagittaria graminea Michaux, which has long-petioled, linear, lanceo- 
late or elliptical leaf blades, acute at both ends, and much smaller flowers 
than S. latifolia, and other less abundant species. A nearly related 
genus is Alisma, represented in our range by the very common Alisma 
subcordatum Rafinesque (American Water Piantain), with oblong, 
elliptic, oval or ovate leaf blades which are cuneate, truncate or cordate at 


38 NEW YORK STATE MUSEUM 


the base, the flowering scapes rather tall, bearing numerous branches and 
pedicels in whorls of three to ten, with very small, white flowers. Like 
most other members of the Alismaceae, it inhabits shallow water or muddy 
places. 
Sedge Family 
Cyperaceae 
Sheathed Cotton Grass or Hare’s Tail Rush 


Eriophorum callithrix Chamisso 
Plate 2a 

The Cotton Grass may be regarded not so much as a wild flower as 
one of the most ornamental of the sedges, since it is not a true grass. It is 
an inhabitant of cold, mossy bogs. The stiff culms, forming tussocks, 
rise eight to twenty inches above the surface of the bog and each culm bears 
at the summit a solitary spikelet of small, perfect flowers; each flower with 
six scalelike divisions, three stamens and a three-cleft style. Within the 
scalelike perianth are numerous soft, white bristles, which become greatly 
elongated in fruit, at which time the bog where the plant is growing becomes 
beautiful with hundreds or thousands of these waving white plumes. 

Common in sphagnum bogs from Newfoundland to Alaska, south to 
Massachusetts, Pennsylvania and Wisconsin. Flowering in early spring, 
the white plumes being at their best in June or, in the far north, in July. 

The Sheathed Cotton Grass is but one of a number of related species 
which add much to the beauty of our wet meadows, swamps and bogs in 
summer. Perhaps even more abundant in the north is the Thin-leaved 
Cotton Grass (Eriophorum viridicarinatum (Engelmann) 
Fernald) with five to thirty nodding white plumes, and the Virginia Cotton 
Grass (Eriophorum virginicum Linnaeus), in which the soft 


bristles of the mature plume are of a dingy brown color. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 1 


BROAD-LEAVED ARROWHEAD 
Sagittaria latifolia 


a 
> 


DUDLULJOADI SIAN 
SSVUD GHAG-MOTIAA VWNITONVO “A 


= 


/ 


L 


XLAY JVI wWindoyd OLAGT 
SSVYD NOLLOD GHHLVAHS “V 


/\ 


| 


| 
'X 


T 938 Id 


MUOA MAN AO SUAMOTA ATIM 


wuMmasnfA 93¥IS *K “N GT Jrowoyyy 


WILD FLOWERS OF NEW YORK 39 


Arum Family 
Araceae 
Jack-in-the-pulpit; Indian Turnip 
Arisaema triphyllum (Linnaeus) Torrey 
Plate 3 

A perennial herb, 1 to 3 feet tall, from a rounded, acrid corm. Leaves 
one or two, nearly erect, and exceeding the scape, three-foliate, the seg- 
ments or leaflets pale green beneath, ovate, acute, rounded or pointed at 
the base, 3 to 8 inches long, 1 to 3 inches wide, unfolding with the flowers. 
Flowers dioecious, borne on the basal part of the club-shaped spadix, which 
is naked, blunt and green or purple above; spathe green and purple-striped, 
curving in a broad flap over the top of the spadix, long pointed, sometimes 
whitish with green stripes or almost uniformly greenish. The crowded 
ovaries of the pistillate flowers ripen into a cluster of bright-red, shining, 
globose berries. 

A common plant of moist woods and thickets, flowering from early 
spring until June. The fruit ripens in July, and in late summer the leaves 
frequently wither and die, leaving the stalks of bright-red berries con- 
spicuous objects in the woods. 

Two closely related species are sometimes recognized, Arisaema 
p usillum (Peck) Nash, with leaves green beneath, a cylindrical spadix 
and spathe deep brown to almost black in color; Arisaema stew- 
ardsonii Britton, with a conspicuously fluted spathe which is whitish 
below and green or green-striped toward the tip, but otherwise resembling 
A. pusillum. 

The Green Dragon or Dragon-root (Arisaema dracontium 
(Linnaeus) Schott) (figure II) has solitary leaves divided into five to seven- 
teen segments, and a narrow greenish or whitish, long-pointed spathe 
enwrapping the spadix, the upper part of which tapers into a slender 
appendage exserted 1 to 7 inches beyond the spathe. The mature berries 
are reddish-orange in color. This plant is less abundant than the Jack-in- 
the-pulpit, and much less conspicuous. 


40 NEW YORK STATE MUSEUM 


Figure IT 
Green Dragon or Dragon-root 


(Arisaema dracontium (Linnaeus) Schott) 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 3 


JACK-IN-THE-PULPIT; INDIAN TURNIP 
Arisaema triphyllum 


Stsnppd DYDD 
WOUV WALVM -VIIVO GTIM 


fF Wid 


si 


MUOA MON AO SHUAMOTA GCTIIM 


winasny 23¥IS “A *N GT owes 


WILD FLOWERS OF NEW YORK 41 


Wild Calla; Water Arum 
Calla palustris Linnaeus 
Plate 4 
A perennial herb of swamps and bogs with long, acrid rootstocks, 
covered with sheathing scales and with fibrous roots at the nodes, from 
which arise numerous petioled leaves with thick, entire, glossy green, 
broadly ovate or suborbicular leaf blades 2 to 5 inches wide, cuspidate or 
pointed at the apex and deeply cordate at the base. Flowering scapes 
about as long as the petioles, sheathed at the base, bearing at the summit 
an ovate-lanceolate or elliptic, acuminate, open spathe, white within and 
greenish without, sometimes with a second spathe nearly opposite the 
first and smaller in size, or rarely the two of equal size. Spadix cylindric, 
much shorter than the spathe, densely covered with perfect flowers, or 
the uppermost flowers staminate. The individual flowers on the spadix 
possess usually six stamens and no perianth. Ovaries ripening into a large 
head of red berries. 
Frequent in swamps and bogs, especially northward. Rare in the 
southern part of the State. Flowering from late May to early July, the 
fruit ripening from June to August. 


Skunk Cabbage 
Spathyema foetida (Linnaeus) Rafinesque 
Plate 5 
A fetid herb, and the first plant to flower in the spring. The leaves 
are large, ovate, cordate, numerous in dense crowns, becoming in summer 
1 to 3 feet long and 1 foot wide, but at flowering time scarcely beginning 
to unfold. Rootstock thick, descending, terminating in whorls of fleshy 
fibers. Spathe preceding the leaves, erect, 3 to 6 inches high, 1 to 3 inches 
in diameter, convolute, firm and fleshy, often one-fourth to one-half of an 
inch thick in the middle, pointed, completely inclosing the spadix, brown 
to greenish yellow, usually mottled, its short scape usually below the surface. 


42 NEW YORK STATE MUSEUM 


Spadix about 1 inch in diameter, entirely covered by the perfect flowers, 
greatly enlarged and sometimes 6 inches in diameter in fruit. The perianth 
of each flower consists of four hooded sepals. 

A common plant of low, wet woods, meadows and swamps. When 
the spathes first appear, they possess little of the rank odor which 
characterizes them when older and which renders them objects of 
opprobrium. They appear almost before the last snowdrifts have dis- 
appeared and indicate the first awakening of plant life in spring. 

The Arum family (Araceae), to which belong the Jack-in-the-pulpit, 
the Wild Calla, and the Skunk Cabbage, also contains several other 
native plants usually found in wet or damp places. The Green Water 
Arum (Peltandra virginica (Linnaeus) Kunth) with bright-green, 
hastate-sagittate leaves, often I to 2 feet long and 3 to 8 inches wide, 
possesses an inconspicuous green spathe, 4 to 8 inches long, with a strongly 
involute undulate margin. The Golden Club (Orontium aquaticum 
Linnaeus), found only in a few localities in the southern part of the State, 
possesses a cylindric, golden yellow spadix, from which the spathe falls 
at flowering time. 

The Sweet Flag, Calamus or Flagroot (Acorus calamus Lin- 
naeus) (figure III) belonging also to this family isa common plant of wet 
meadows, with long, linear, flaglike leaves and the spathe a leaflike exten- 
sion of the scape, the spadix spikelike, 2 to 3 inches long and about 
one-half of an inch in diameter, compactly covered with minute greenish 


yellow flowers. 
Yellow-eyed Grass Family 


Xyridaceae 
Carolina Yellow-eyed Grass 
Xyris caroliniana Walter 
Plate 2b 
A small, tufted, grasslike plant of wet meadows and bogs, with 
numerous fibrous roots and flat, linear, grasslike leaves 4 to 15 inches long. 
Flowering scapes as long or usually much longer than the leaves. Some- 


times over a foot tall, bearing at the summit a dense, ovoid, obtuse spike 


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Sweet Flag, Calamus or Flagroot 


(Acorus calamus Linnaeus) 


44 NEW YORK STATE MUSEUM 


of coriaceous, overlapping bracts or scales. Flowers bright yellow, about 
one-fourth of an inch broad, on short, slender peduncles; each flower con- 
sisting of three oblong divisions to the corolla, three stamens inserted on 
the corolla and a three-branched style. Flowers appearing from the axils 
of the bracts comprising the spike. 

In swamps, bogs and borders of streams from Maine to Florida and 
Louisiana, mostly near the coast. Flowering from June to August; in 
New York usually flowering in July. 

Several closely related species are recognized by botanists. XKyris 
flexuosa Muhlenberg, of about the same range, has the scapes bulbous- 
thickened at the base. Xyris montana H. Ries, in bogs from Nova 
Scotia to Michigan and Pennsylvania, is frequent in the mountainous 
parts of New York, and resembles very closely the Carolina Yellow-eyed 
Grass. 

Pipewort Family 
Bit tocadwliace are 
Seven-angled Pipewort 
Eriocaulon septangulare Withering 
Plate 6a 

A small, tufted plant with pellucid, fenestrate-nerved leaves arising 
from the crown, I to 3 inches long. Flowering scapes weak, twisted, 
usually about seven-angled, 1 to 8 inches tall, or when submersed, both 
the leaves and the scapes considerably elongated, bearing at the summit 
a small subglobose head of woolly white flowers, interspersed with numerous 
bracts; most of the flowers staminate and about one-eighth of an inch 
long, the few pistillate flowers scarcely more than half as large. 

In still water and on shores of ponds, lakes and streams, usually where 
it is sandy, from Newfoundland to Minnesota, Florida and Texas. Usually 
in flower in this State in July and August. In the south are numerous 


other species of this and related genera. 


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WILD FLOWERS OF NEW YORK 45 


Spiderwort Family 
Commelinaceae 
Asiatic Dayflower 
Commelina communis Linnaeus 
Plate 6b 

A glabrous plant with decumbent or ascending branching stems, often 
rooting at the nodes, 1 to 3 feet long with numerous oblong-lanceolate 
leaves 3 to 5 inches long and 1 to 13 inches wide, acuminate at the tips 
and narrowed or rounded at the base, smooth and dark green, the stem 
with white-membranous, green-veined sheaths below each leaf. Flowers 
toward the ends of the branches or stems, each subtended by green leaflike 
spathes about 1 inch long, deep blue, one-half of an inch broad or broader, 
irregular, consisting of three sepals and three petals, two of them much 
larger than the third; three fertile stamens, one of them incurved and 
its anther larger than the others. In addition there are three sterile stamens. 
Fruit a small, two-celled capsule, each cell with two seeds; seeds compressed, 
dark brown and roughened. 

A native of Asia, commonly naturalized or adventive in southern New 
York and southward, and occasionally appearing farther north. Sometimes 
called “‘ Wandering Jew.”’ 


Spider Lily; Spiderwort 
Tradescantia virginiana Linnaeus 
Plate 7 

A tall, smooth or slightly pubescent plant, belonging to the same 
family as the Dayflower, often 1 to 3 feet tall, with long, linear or linear- 
lanceolate, long-pointed leaves, often a foot long and one-half to 1 inch 
wide, usually more or less channeled along the middle. Flowers in terminal 
umbels or clusters on slender pedicels, one-half to 2 inches long; blue or 
purplish, rarely white, 1 to 2 inches broad with three small sepals, three 
large, obovate, similar petals, and six equal and fertile stamens. 


46 NEW YORK STATE MUSEUM 


In rich soil, mostly in woods and thickets, from southern New York, 
Ohio and South Dakota, south to Virginia and Arkansas. A common 
plant in cultivation farther north where it is a frequent escape to roadsides 
and fence rows. Although of great beauty, the flowers are of brief duration, 
and the delicate petals soon wither, the flowers being followed by others 
until all the numerous buds of each cluster have bloomed. With us it 
usually flowers in June and July’or sometimes as late as August, especially 
since not all the stems appear to reach maturity at the same time. 


Pickerel Weed Family 
Pontederiaceae 
Pickerel Weed 
Pontederia cordata Linnaeus 
Plate 8 

A perennial aquatic herb, rising from a thick, horizontal rootstock, 
with thick, glossy, dark-green, ovate to lanceolate leaves, cordate-sagittate, 
truncate or narrowed at the base, 2 to 10 inches long, I to 6 inches wide, 
the apex and basal lobes obtuse. Flowering stems erect, 1 to 4 feet tall, 
glandular-pubescent above, one-leaved, with several sheathing, bractlike 
leaves at the base. Flowers blue, ephemeral, numerous, in a dense head 
or spike (spadix) subtended by a thin bractlike spathe. Each flower is 
tubular, about one-fourth of an inch long, curved, two-lipped, the upper 
lip composed of three ovate lobes, of which the middle one is the longest, 
and with two yellow spots at the base within, the lower lip of three linear- 
oblong spreading lobes. Stamens six, the filaments, anthers and style 
bright blue. After flowering, the lobes and upper part of the perianth tube 
wither above, while the persistent base hardens around the fruit. 

Frequent along the borders of ponds and streams and shallow margins 
of lakes, where it flowers from June to September, usually at its best in 
August. One of the most attractive of our native aquatic plants. 

The Pickerel Weed belongs to the Pontederiaceae, represented in 
our flora by but one other genus, the Mud Plantains (Heteranthera), with 


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WILD FLOWERS OF NEW YORK 47 


two species of small, inconspicuous herbs of shallow water or wet muddy 
shores. 
Bunchflower Family 


Melanthaceae 


Glutinous Triantha or False Asphodel 
Triantha glutinosa (Michaux) Baker 
Plate ob 

A perennial, herbaceous plant, somewhat bulbous at the base; stems 
6 to 20 inches high, viscid pubescent with black glands, bearing a few leaves 
near the base; most of the leaves basal and tufted, linear and grasslike, 
2 to 7 inches long. Flowers numerous in an oblong raceme at apex of the 
stem, subtended by minute involucral bracts, each flower about one-fourth 
of an inch broad, the perianth white, divided into six nearly equal oblong 
segments; stamens six, with pink or reddish anthers; fruit a small oblong 
capsule, the tiny seeds with a curved appendage at each end. 

An inhabitant of sphagnum or marly bogs from Newfoundland to 
Minnesota, Michigan and the southern Alleghanies. By no means a 
common plant, and one which the wild flower connoisseur always likes to 
locate, and which repays by its rarity rather than its beauty the inevitable 
journey to the boggy place where it grows. 


Glaucous Anticlea 
Anticlea chlorantha (Richardson) Rydberg 


Plate 9a 

A slender, herbaceous, perennial plant from a membranous coated, 
ovoid bulb which is about an inch long. Stems slender, 6 inches to 3 feet 
tall; leaves linear, one-eighth to seven-eighths of an inch wide, keeled, the 
lower ones 4 to 12 inches long, the upper ones much shorter. Inflor- 
escence a simple, open raceme or large, loose panicle, 4 to 12 inches long, 
with slender, ascending branches. Flowers perfect, greenish or yellowish, 
about three-fourths of an inch broad; perianth segments oval or obovate, 
obtuse, bearing a large obcordate gland just above the short claw, the 


45 NEW YORK STATE MUSEUM 


perianth persistent and adnate to the lower part of the ovary after withering. 
Fruit an oblong, three-celled capsule, about 1 inch long. 

An inhabitant of marshes, bogs and moist places, New Brunswick to 
Vermont, New York, Manitoba and Missouri. Flowering in August. 
Not so rare as the Glutinous Triantha, but usually growing in similar 
locations. 

Bunchflower 
Melanthium virginicum Linnaeus 
Plate 10 

A rather tall, leafy, herbaceous plant, perennial by a thick rootstock; 
stems slender to somewhat stout, 2 to 5 feet high. Leaves linear, acumi- 
nate, often a foot long, but only one-third to 1 inch wide, the lower ones 
sheathing the stem, the upper ones smaller and sessile. Inflorescence a 
many-flowered panicle, 6 to 18 inches long, pubescent; flowers about three- 
fourths of an inch broad, greenish yellow, turning brown with age; perianth 
of six spreading, separate, persistent segments, each segment consisting of 
an oblong, obtuse, flat blade, sometimes obcordate, about twice as long as 
the claw, and bearing two dark glands at its base; stamens shorter than 
the segments and adnate to them; fruit a three-lobed capsule, about two- 
thirds of an inch long. 

In meadows, wet woods and marshes, Rhode Island to southern New 
York and Minnesota, south to Florida and Texas. Flowering in July 
and August. 

The Bunchflower is not a common plant in New York, and is found 
only in a few localities in the southern part of the State. Two closely 
related species are equally uncommon and also restricted to the southern 
part of the State. These are: (1) the Crisped or Broad-leaved Bunch- 
flower (Melanthium latifolium Desvaux), with broader leaves 
and the blade of the perianth segments undulate, crisped and scarcely 
longer than the claw; (2) the Pine-barren Oceanorus (Oceanorus 
leimanthoides (A. Gray) Small), with linear, blunt leaves, the outer 


ones becoming fibrous, and small whitish flowers with oblong perianth 
segments, which are sessile. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum _ Plate 9 x 
c on 


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A. GLAUCOUS ANTICLEA B. GLUTINOUS TRIANTHA; FALSE ASPHODEL 
Anticlea chlorantha Triantha glutinosa 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 10 


BUNCHFLOWER 
Melanthium virginicum 


WILD FLOWERS OF NEW YORK 49 


American White Hellebore; Indian Poke 
Veratrum viride Linnaeus 
Plate 11 

A tall, leafy, perennial herb; stem simple and branched only in the 
inflorescence, stout, 2 to 8 feet tall, from a poisonous, perennial, erect root- 
stock 2 to 4 inches long and 1 to 2 inches thick with numerous fleshy-fibrous 
roots. Leaves alternate, clasping, strongly veined and plaited, all pointed 
at the apex; the lower leaves broadly oval or elliptic, 5 to 12 inches long, 
3 to 6 inches wide; the upper ones successively narrower and shorter, those 
of the inflorescence very small; stem and inflorescence pubescent. Flowers 
greenish or greenish yellow, two-thirds to 1 inch broad arranged in a many- 
flowered panicle at the summit of the stem, the panicle often 8 to 20 inches 
long, its lower branches spreading or drooping. Segments of the perianth 
six, without glands or claws, oblong or oblanceolate, ciliate-serrulate, twice 
as long as the curved, yellow stamens which are six in number and opposite 
the perianth segments. Fruit a three-lobed, three-celled, many-seeded 
capsule, three-fourths to 1 inch long and one-third to one-half of an 
inch thick. 

In swamps and wet woods, New Brunswick, Quebec and Ontario to 
Minnesota, south to Georgia and Tennessee. Flowering in May and June, 
or July in the far north. 

Lily Family 
Ie eli arcierare 
Day Lily 
Hemerocallis fulua Linnaeus 
Plate 12 

A tall, glabrous herb with fibrous roots, usually growing in dense 
clusters. Leaves mostly basal, linear, erect or spreading, 1 to 2 feet long 
and one-half to two-thirds of an inch wide, channeled and tapering to 
an acute tip. Flowering scapes leafless, 2 to 5 feet tall, bearing a few 
short bracts above, and six to fifteen flowers on short pedicels. Perianth 


50 NEW YORK STATE MUSEUM 


funnelform, tawny-orange, 4 to 5 inches long, opening for a day, its lobes 
oblong, somewhat spreading, netted-veined, the three outer nearly flat 


and more acute; the three inner ones undulate on their margins and blunt. 
Stamens six, inserted at the top of the perianth tube, shorter than the 


lobes of the perianth and declined. Fruit an oblong, thick-walled, three- 
angled, wrinkled capsule. 

Native of Europe and Asia. Frequent in cultivation in this country 
and commonly escaped to meadows, along streams and roadsides or 
persistent in old yards and cemeteries. A flower of early introduction 
into the eastern states where it has made itself quite at home as an 
escape, and thrives and spreads with amazing rapidity under favorable 
conditions. 

The Yellow Day Lily (Hemerocallis flava Linnaeus), with 
yellow flowers, their lobes parallel-veined, is occasionally found near old 
gardens and on roadsides, but not so frequently as the tawny-orange flowered 
Day Lily. 

Red Lily; Wood Lily; Philadelphia Lily 
Lilium philadelphicum Linnaeus 
Plate 13 

Stems 1 to 3 feet tall from a bulb about I inch in diameter and com- 
posed of numerous narrow, jointed, fleshy scales. Leaves lanceolate, 
acute at both ends, or the lower leaves sometimes obtuse, in whorls of 
three to eight on the stem, or a few of the upper leaves alternate, thin, with 
finely roughened margins. Flowers one to five at summit of stem, erect, 
3 to 4 inches high; perianth reddish orange, its six equal segments spatulate, 
somewhat spreading, pointed or obtuse, one-half to 1 inch wide, gradually 
narrowed below, spotted with purple toward the base; stamens six, about 
as long as the club-shaped style. Capsule oblong-ovoid, 1 to 2 inches 
long. 

In rather dry woods and thickets, more often in sandy regions than 
elsewhere, Maine and Ontario to North Carolina and West Virginia. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 11 


AMERICAN WHITE HELLEBORE; INDIAN POKE 
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Memoir 15 N. Y. State Museum Plate 13 


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RED, WOOD OR PHILADELPHIA LILY 
Lilium philadelphicum 


WILD FLOWERS OF NEW YORK 51 


Wild Yellow Lily; Canada or Nodding Lily 
Lilium canadense Linnaeus 
Plate 14 

Stems 2 to 5 feet tall, from a stout rootstock bearing several subglobose, 
scaly, white bulbs. Leaves in whorls of four to ten or some of them alter- 
nate, lanceolate or oblong-lanceolate, acuminate, 2 to 6 inches long, one- 
fourth to 1% inches wide, finely roughened on the margins and on the 
veins beneath. Flowers one to sixteen, nodding on long peduncles at the 
top of the stem; perianth segments 2 to 3 inches long, yellow or red, usually 
thickly spotted, recurved or spreading; fruit an oblong, erect capsule 1 to 2 
inches long. 

Common in swamps, moist meadows, and fields, Nova Scotia to 
Minnesota, Georgia, Alabama and Nebraska. Flowering in July and 
August. A common and most attractive wild flower of the east, more 
abundant than the Turk’s-cap Lily (Lilium superbum Linnaeus), 
which has similar but usually larger flowers, usually orange-red and purple- 
spotted, more strongly recurved flower segments and leaves smooth and 
not roughened on the margins or veins as in L. canadense. 


Yellow Adder’s-tongue; Dog’s-tooth Violet 
Erythronium americanum Ker 
Plate 15a 

A low, herbaceous plant arising from a deeply buried corm which 
propagates by offshoots; the simple stem 6 to 12 inches long, bearing a pair 
of equal or somewhat unequal, oblong or oblong-lanceolate, flat leaves, 
dark glossy green, usually mottled with brown, sometimes green all over, 
narrowed into clasping petioles; the flower stem arising from between 
the leaves, bearing a single nodding flower; perianth yellow or rarely 
purplish-tinged, the segments oblong, seven-eighths to 2 inches long, about 
one-fourth of an inch wide or less, recurved, dotted within, the three inner 
ones auricled at the base; style club-shaped; capsule obovoid. 


NEW YORK STATE MUSEUM 


on 
Ny 


In moist woods and thickets, Nova Scotia to Ontario and Minnesota 
south to Florida and Arkansas. Flowering in April and May. 

One of the commonest and best known of our spring flowers, coming 
in company with the Hepatica, Spring Beauty, and Squirrel Corn, but 
usually in its prime a little later than these. The white Dog’s-tooth Violet 
(Erythronium albidum Nuttall), with leaves less or not at all 


spotted and pinkish white flowers, is very rare. 


Ague or Colicroot; Star Grass 
Aletris farinosa Linnaeus 
Plate 16 

Leaves mostly basal, lanceolate and spreading, forming a dense cluster, 
lanceolate, long pointed, narrowed at the base, pale yellowish green, 2 to 
7 inches long, one-fourth to 1 inch wide. Roots numerous, tough and 
very bitter. Stem or scape 1 to 3 feet tall, bearing a few distant bractlike 
leaves. The terminal raceme of flowers 4 to 12 inches long; flowers erect 
on short pedicels subtended by small bracts; perianth tubular-oblong, six- 
lobed, white or the short lobes yellowish, about one-fourth to one-third 
of an inch long and less than half as thick, mealy-roughened without; 
capsules ovoid, about one-sixth of an inch long, inclosed by the withering- 
persistent perianth. 

In dry, mostly sandy soil, Maine to Ontario and Minnesota, south to 
Florida and Arkansas. Flowering in June and July. In New York rarely 
seen except in the sandy regions adjacent to the coast. Extremely abun- 
dant on sterile sandy fields like the Hempstead plains of Long Island, where 
it is very conspicuous in early summer. It possesses a number of 
vernacular names, such as Ague Grass, Blazing Star, Bitter Grass, Crow 
Corn, Mealy Starwort, Aloeroot, Starroot, Huskroot, and others. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 14 


WILD YELLOW LILY; CANADA OR NODDING LILY 
Lilium canadense 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 15 


A. YELLOW ADDER’S-TONGUE; DOG’S-TOOTH VIOLET 
Erythronium americanum 


B. CLIMBING FALSE BUCKWHEAT 
Bylderdykia scandens 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 


Plate 16 
ees) 


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AGUE OR COLICROOT; STAR GRASS 
Aletris farinosa 


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WILD FLOWERS OF NEW YORK 53 


Lily of the Valley Family 
Convallariaceae 
Yellow Clintonia; Dogberry 
Clintonia borealis (Aiton) Rafinesque 
Plate 17 

Flowering scape or stem 6 to 15 inches high, with two to five (usually 
three), oval, oblong or obovate, thin, glossy green leaves at the base, their 
petioles sheathing the base of the stem which arises from a slender root- 
stock. Leaves ciliate, 5 to 8 inches long, and 14 to 34 inches wide. Flowers 
three to six, forming an umbel at the top of the stem, sometimes a secondary 
cluster of flowers below the top, drooping, greenish yellow, three-fourths 


to 1 inch long, on pedicels about as long as the flowers; perianth segments 


g; 
distinct, six in number, equal and somewhat spreading, the six stamens 
about as long as the perianth. Fruit an oval, dark blue, shining berry 
about one-fourth of an inch in diameter. 

In moist woods and thickets, Newfoundland to Manitoba, south to 
North Carolina and Wisconsin. Very common in the rich, moist wood- 
lands of northern New York, but rare or absent from the coastal region. 
Flowering from the latter part of May until the last of June. The fruit 
ripe in September. 

The White Clintonia (Clintonia umbellulata (Michaux) 
Torrey), with smaller white flowers, not drooping, and black berries, leaves 
and scapes more pubescent but otherwise similar, which is common in the 
southern Appalachians, reaches New York in the southwestern counties 
of the State. 

Wild or False Spikenard; False Solomon’s-seal 
Vagnera racemosa (Linnaeus) Morong 
Plate 18 

Stem slender or stout, erect or ascending, sometimes zigzag, 1 to 3 feet 
tall, simple, bearing numerous alternate, sessile or nearly sessile oblong- 
lanceolate or oval, acuminate leaves, 3 to 6 inches long, I to 3 inches wide, 


54 NEW YORK STATE MUSEUM 


which are finely pubescent, especially beneath, and sometimes also above, 
their margins minutely ciliate. Rootstock rather thick and fleshy with 
numerous long, fibrous roots, the scars of former stems irregular and ring- 
like. Flowers white; many, forming a large terminal panicle, 1 to 4 inches 
long; each flower about 2 lines broad; perianth of six oblong, equal, separate, 
spreading segments. Fruit a red, aromatic berry about 3 lines in diameter 
and speckled with purple. In Bergen swamp occurs a variety with three 
to six purple stripes like the fruit of V. stellata. 

In moist woods and thickets, Nova Scotia to British Columbia, south 
to Georgia, Missouri and Arizona. 

The Star-flowered Solomon’s-seal (Vagnera stellata (Linnaeus) 
Morong) is scarcely less abundant, but seems to prefer thickets and banks 
with more moisture. The leaves are smaller and narrower, the flowers 
fewer in number, larger, white, and racemed; the berries green with six 
black stripes, or entirely black. 


Three-leaved Solomon’s-seal 
Vagnera trifolia (Linnaeus) Morong 
Plate 32a 

Stem and leaves glabrous from a slender, elongated rootstock, the erect 
stem 2 to 15 inches high with two to four (usually three) oval, oblong or 
oblong-lanceolate, sessile leaves 2 to 5 inches long, one-half to 2 inches 
wide with sheathing bases. Flowers white, few, racemed at the top of the 
stem; perianth segments oblong or oblong-lanceolate, obtuse, finally some- 
what reflexed and longer than the stamens. Fruit a dark-red berry about 
one-fourth of an inch in diameter. ; 

In bogs and wet woods, Newfoundland to British Columbia, south to 
Connecticut, New Jersey, Pennsylvania and Michigan. Usually abundant 
in sphagnum under or near spruces and tamaracks, and therefore rare out- 
side the mountainous sections of the State except on the margins of bogs 
and mossy swamps where the spruce and tamarack abound, as, for instance, 
Cicero swamp in Onondaga county, and Bergen swamp in Genesee county, as 


well as numerous other swamps of similar character throughout the State. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 17 


YELLOW CLINTONIA; DOGBERRY 
Clintonia borealis 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 18 


WILD OR FALSE SPIKENARD; FALSE SOLOMON’S-SEAL 
Vagnera racemosa 


WILD FLOWERS OF NEW YORK 55 


False or Wild Lily of the Valley 
Two-leaved Solomon’s-seal 
Unifolium canadense (Desfontaines) Greene 
Plate 19 

A low, herbaceous perennial with slender rootstock, and _ slender, 
erect and often zigzag stem, 2 to 7 inches high, bearing one to three (usually 
two) ovate or ovate-lanceolate, pointed leaves, cordate at the base and 
sessile or short-petioled; stemless plants frequent and consisting of a 
single leaf on a petiole 1 to 4 inches long arising from the rootstock. Flowers 
white, numerous, forming a rather dense terminal raceme, I to 2 inches 
long; perianth about one-fifth of an inch broad with four spreading, 
separate segments, which slightly exceed the four stamens in length; fruit 
a cluster of pale-red, speckled berries, each with one or two seeds. 

In moist woods and thickets, Newfoundland to the Northwest Territory 
of Canada, south to the mountains of North Carolina, Tennessee, Iowa 
and South Dakota. Flowering in May and June. 

A common wild flower of most parts of New York, especially in the 
moist cool forests of the northern and mountainous sections. 


Sessile-leaved Bellwort 
Uvularia sessilifolia Linnaeus 
Plate 20a 

Stems slender, glabrous, 4 to 12 inches high, naked or with one or 
two leaves below the fork. Leaves oblong or oblong-lanceolate, 1 to 3 
inches long when they mature, thin, sessile, acute at each end, slightly 
rough-margined, pale or glaucous beneath; flowers greenish yellow, two- 
thirds to 11 inches long; the six perianth segments smooth, the stamens 
shorter than the styles; anthers blunt; fruit a sharply three-angled capsule, 
narrowed at both ends, about 1 inch long and two-thirds as thick. 

In moist woods and thickets, usually most abundant where the soil 
is sandy, New Brunswick and Ontario to Minnesota, south to Georgia 


56 NEW YORK STATE MUSEUM 


and Arkansas. Except in the coastal region and other sections of the 
State with sandy soil, this small-flowered bellwort is not so abundant 
asU. grandiflora and U. perfoliata. 


Large-flowered Bellwort 
Uvularia grandiflora J. E. Smith 
Plate 20b 

An erect herb with smooth, leafy, forked stem from a perennial root- 
stock; stem with one or two leaves below the fork, 6 to 20 inches high. 
Leaves perfoliate, oblong, oval or ovate, pubescent beneath, glabrous above, 
becoming 2 to 5 inches long, acute at the apex, rather smaller and often 
scarcely unfolded at flowering time; flowers solitary at the ends of the 
branches, peduncled, drooping, narrowly bell-shaped, lemon-yellow, I to 13 
inches long, perianth segments six, distinct, smooth on both sides or very 
slightly granular within; stamens six, longer than the styles which are 
united to about the middle; anthers linear, the connective blunt; fruit a 
three-angled, truncate capsule, about one-half of an inch long. 

In rich upland woods, Quebec to Ontario, Minnesota, Georgia and 
Kansas. Flowering in April and May. A common flower of most sections, 
especially in rich woodlands. In the Ontario lowlands and Hudson valley 
it is largely replaced by the Perfoliate Bellwort (Uvularia per- 
foliata Linnaeus), which differs chiefly in having smooth and glaucous 
foliage, and slightly smaller flowers with the perianth segments papillose 
within. 

Sessile-leaved Twisted-stalk 
Streptopus roseus Michaux 
Figure IV 

Stems 1 to 3 feet high from a short, stout rootstock covered with fibrous 
roots. Branches usually three or four in number and obliquely ascending, 
all leafy and sparingly pubescent. Leaves alternate, thin, many-nerved, 
ovate or ovate-lanceolate, 2 to 44 inches long, long pointed at the apex, 
sessile, rounded or slightly clasping the stem at the base, green on both 


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WILD FLOWERS OF NEW YORK SH 


sides, but usually paler beneath, their margins finely ciliate. Flowers 
purple or rose colored, about one-third of an inch long,  slender- 
peduncled, solitary or two together from the axil of each of the upper leaves 
on slender peduncles, one-half to 1 inch long, the peduncle bent or twisted 
at about the middle; perianth bell-shaped, its six segments lanceolate and 
pointed, their tips somewhat recurved or spreading. Stamens six, shorter 
than the perianth. Fruit a globose, red berry about one-third of an inch 
or less in diameter. 

In moist woods, Newfoundland to Manitoba, Georgia and Michigan. 
Flowering in May and June. 


The Clasping-leaved Twisted-stalk (Streptopus amplexi- 
folius (Linnaeus) De Candolle) is similar but the leaves are clasping 
around the stem at their bases, glaucous or whitish beneath and the flowers 
are greenish white in color. 

The Hairy Disporum (Disporum lanuginosum (Michaux) 
Nichols.) resembles the Twisted-stalks in manner of growth, but the leaves 
are somewhat narrower and not clasping and the flowers are solitary or 
few together at the ends of the branches, one-half to three-fourths of an 
inch long and greenish in color. It is found in woods in western New York. 


Hairy Solomon’s-seal 
Polygonatum biflorum (Walter) Elliott 


Plate 21 


Stems slender, smooth, arching, often zigzag above from a thick, 
horizontal, jointed rootstock, bearing the raised orbicular scars of the stems 
of former years; stem naked below, above bearing six to many opposite 
or nearly opposite, oval or ovate leaves, 2 to 4 inches long, one-half to 2 
inches wide, acute or acuminate at the apex, narrowed or obtuse at the 
base, pale or pubescent beneath, glabrous above, the upper leaves commonly 
narrower than the lower; flowers in drooping, axillary clusters of one to 
four (often two), perianth greenish or greenish yellow, tubular, one-third 
to one-half of an inch long, with six short lobes; the six stamens shorter 
than the tube, their anthers sagittate and filaments minutely roughened. 
Fruit a dark blue, pulpy berry about one-fourth of an inch in diameter. 


58 NEW YORK STATE MUSEUM 


Figure IV 
Sessile-leaved Twisted-stalk 


(Streptopus roseus Michaux) 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 21 


le: : 


HAIRY SOLOMON’S-SEAL 
Polygonatum biflorum 


WILD FLOWERS OF NEW YORK 59 


A common but not showy plant of woods and thickets from 
New Brunswick to Ontario and Michigan, south to Florida and 
Tennessee. 

Resembling this but usually larger in every way, with glabrous 
leaves and smooth filaments, is the Smooth or Giant Solomon’s-seal 
(Polygonatum commutatum (Roemer & Schultes) Dietrich), 
which ranges northeastward only to Rhode Island, New Hampshire and 
Ontario. It seems to prefer moist thickets and woods along streams and 
on bottomlands, while the Hairy Solomon’s-seal is more commonly met 
with in rich upland woods. 


Wake-robin Family 
iMeawlilttarcerare 
Indian Cucumber Root 
Medeola virginiana Linnaeus 
Plate 22 
A slender, erect, unbranched herb from a perennial rootstock 1 to 3 
inches long; stem 1 to 23 feet high, loosely covered with deciduous wool, 
bearing the lower whorl of leaves above the middle or, in flowerless plants, 
at the summit; leaves of the lower whorl sessile, 2 to 5 inches long, 1 to 2 
inches wide; acuminate at the apex, narrowed at the base, three to five- 
nerved; leaves of the upper whorl 1 to 2 inches long, one-half to 1 inch 
wide, short petioled or sessile, often turning reddish at the base; umbel 
of two to nine flowers on filiform pedicels, 1 inch long or less, declined in 
flower, erect or ascending in fruit; perianth segments one-fourth to one- 
half of an inch long, obtuse, the six equal segments recurved, the three 
long styles recurved. Fruit a dark blue or purplish berry one-fourth 
to one-half of an inch in diameter. 
In moist woods and thickets, Nova Scotia to Ontario, Minnesota, 
Florida and Tennessee. Flowering in May and June. Fruit ripe in 


September. 


60 NEW YORK STATE MUSEUM 


The thick, tuberlike, white rootstock is brittle with numerous slender 
fibrous roots and has the odor and taste of cucumbers. 


Red Trillium; Wake-robin or Birthroot 


Trillium erectum Linnaeus 


Plate 23a 

Stem rather stout, 8 to 16 inches high, from a thick, short rootstock. 
Leaves rather dark green, very broadly rhombic, 3 to 7 inches long, often 
as wide or wider, sessile or nearly so, acuminate at the apex, narrowed 
at the base, peduncle 1 to 4 inches long, erect or nearly so, bearing a single, 
unpleasantly scented, large flower; sepals lanceolate, acuminate, spreading, 
one-half to 14 inches long; petals lanceolate to ovate, acute, spreading, 
equalling the sepals or a little longer, dark purplish-red, varying to pink; 
greenish, white, or reddish yellow in certain aberrant forms; anthers longer 
than the filaments and exceeding the stigmas; ovary purple with short- 
spreading or recurved styles; fruit an ovoid, somewhat six-lobed, reddish 
berry, I inch thick or less. 

In woods and thickets, Nova Scotia to Ontario, south to North 
Carolina and Tennessee. 


White Trillium; Large-flowered Wake-robin 
Trillium grandiflorum (Michaux) Salisbury 
Plate 23b 

A glabrous, erect, unbranched herb from a stout, perennial, short, 
scarred rootstock, 8 to 18 inches high; bearing at the top of the stem three 
light-green, broadly rhombic-ovate or rhombic-oval leaves, 2 to 6 inches 
long, acuminate at the apex, narrowed and sessile at the base, peduncle 
erect or nearly so, I to 3 inches long, bearing a single flower 2 to 3 inches 
broad. The three sepals lanceolate, pointed and spreading. Petals three, 
erect-spreading, oblanceolate, obovate, or rarely ovate-oblong, obtuse or 
cuspidate, thin, strongly veined, white, usually turning pink with age, 
much longer than the sepals. Stamens six, with yellow anthers which are 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum : Plate 22 


INDIAN CUCUMBER ROOT 
Medeola virginiana 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 23 


A. RED TRILLIUM; WAKE-ROBIN OR BIRTHROOT 
Trillium erectum 


B. WHITE TRILLIUM; LARGE-FLOWERED WAKE-ROBIN 
Trillium grandiflorum 


WILD FLOWERS OF NEW YORK 61 


about one-half of an inch long; the three styles slender and ascending. 
Fruit a globose, black, slightly six-lobed berry, three-fourths to 1 inch in 
diameter. 

In rich woods, preferring ravines and wooded upland slopes, Quebec 
to Ontario and Minnesota, south to North Carolina and Missouri. 

Various monstrous forms sometimes occur with two to several long- 
petioled leaves, double flowers, and even forms with green, variegated or 
leaflike petals. 

Painted Wake-robin 


Trillium undulatum Willdenow 
Plate 24a 

Stem slender, 8 to 20 inches high, bearing three ovate, petioled, bluish 
green, waxy leaves, 3 to 8 inches long, 2 to 5 inches wide, long-acuminate 
at the apex, obtuse or rounded at the base. Flowers on erect or somewhat 
inclined peduncles, 1 to 23 inches long; sepals lanceolate, acuminate, three- 
fourths to 14 inches long, spreading; petals ovate or ovate-lanceolate, 
acuminate or acute, white and marked with magenta veins at the base, 
thin, and longer than the sepals, widely spreading, wavy-margined; the 
ovoid ovary with three slender spreading styles, ripening into an ovoid, 

obtuse, bluntly three-angled bright red shining berry. 
Common in woods, especially low, moist or cool, sandy woodlands, 
Nova Scotia to Ontario and Wisconsin, south to Georgia and Missouri. 
Flowers in May and usually a few days later than the white or red trilliums. 


Nodding Wake-robin 
Trillium cernuum Linnaeus 
Plate 24b 
Stems rather slender, 8 to 20 inches high; leaves pale green, broadly 
rhombic, acuminate at the apex, narrowed at the base, sessile or with very 
short petioles, peduncle one-half to 13 inches long, recurved beneath the 
leaves and bearing a single drooping flower about 1 to 14 inches broad, 
sepals lanceolate or ovate-lanceolate, acuminate, petals white or pinkish, 


62 NEW YORK STATE MUSEUM 


ovate-lanceolate or oblong-lanceolate, rolled backward, wavy-margined, 
two-thirds to 1 inch long, as long or longer than the sepals; anthers about 
as long as the subulate filaments and overtopping the stout, recurved styles, 
ovary whitish, ripening into an ovoid reddish-purple berry. 

In rich, usually low woodlands Newfoundland to Ontario and 
Manitoba, south to Georgia and Missouri. Flowering in May or in the 
extreme northern part of its range in June, usually about ten days later 
than the white or red trilliums. 


Amaryllis Family 
Amaryllidaceae 
Yellow Star Grass 
Hypoxis hirsuta (Linnaeus) Coville 
Plate 25 

A low, perennial herb with the leaves all basal, narrowly linear, one- 
eighth to one-fourth of an inch wide, more or less villous and mostly 
longer than the flowering stems, from an ovoid or globose corm, one-fourth 
to one-half of an inch in diameter; flowering stems erect, slender, villous 
above, usually glabrous below, 2 to 6 inches high, bearing at the summit 
an umbel of one to seven flowers; perianth six-parted, its segments narrowly 
oblong, spreading, equal or nearly so, obtuse, bright yellow within, greenish 
and villous without, one-fourth to one-half of an inch long; stamens some- 
what unequal; the style rather shorter than the stamens and three-angled, 
the stigma decurrent on the angles; capsules in fruit about one-eighth 
of an inch in diameter, the black seeds angled. 

In dry, especially sandy soil, in fields, thickets and open woods, Maine 
to Ontario, Assiniboia, Kansas, Florida and Texas. Flowering in May 
and June, but frequently putting up additional flowering scapes as late 
as August and September. 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 7. Plate 25 


YELLOW STAR GRASS 
Hypoxis hirsuta 


WILD FLOWERS OF NEW YORK 63 


Iris Family 
Iridaceae 
Larger Blue Flag 
Iris versicolor Linnaeus 
Plate 26 
Stems round and smooth, erect, sometimes flexuous, 2 to 3 feet tall, 
often branched above, leafy; leaves erect, shorter than and chiefly upon 
the lower part of the stem, somewhat glaucous, one-half to 1+ inches 
wide; rootstock horizontal, thick, fleshy, covered with the fibrous roots. 
Flowers several, the perianth consisting of six, clawed segments united 
below into a tube, the three outer ones dilated, reflexed, violet-blue, varie- 
gated with yellow, green and white; crestless, spatulate, 2 to 3 inches long, 
and wider and longer than the three inner segments; the ovary below the 
perianth tube, in fruit becoming an oblong, obscurely three-lobed capsule, 
I to 1% inches long; divisions of the style petallike, arching over the 
stamens, bearing the stigmas immediately under their two-lobed tips. 
In marshes, thickets and wet meadows, common along streams and 
ponds, Newfoundland to Manitoba, south to Florida and Arkansas. 


Narrow Blue Flag; Poison Flagroot 


Iris prismatica Pursh 
Plate 27 

More slender in every way than Iris versicolor Linnaeus, 
with a tuberous-thickened rootstock; stem 1 to 3 feet tall, bearing two 
or three very narrow, almost grasslike leaves usually less than one-fourth 
of an inch wide; flowers one or two at summit of each stem, blue, veined 
with yellow on slender pedicels; outer perianth segments one-half to 2 
inches long, smooth and devoid of a crest, the inner segments smaller 
and narrower; the perianth tube about one-fourth of an inch long above 
the ovary. Fruit a narrowly oblong capsule, acute at each end and sharply 
three-angled, 1 to 13 inches long. 


64 NEW YORK STATE MUSEUM 


Marshes, wet meadows and swamps, mainly near the coast from Nova 
Scotia to Pennsylvania and Georgia. Flowering in May and June. 


Pointed Blue-eyed Grass 
Sisyrinchium angustifolium Miller 
Plate 28a 

Perennial and tufted, stems stiff and erect, pale green and glaucous, 
4 to 18 inches high. Leaves pointed, about half as long as the stems, 
about one-sixth of an inch or less wide. Stem simple or rarely branched, 
winged, the edges minutely serrulate. Flowers deep violet-blue, one-half 
of an inch broad, umbellate from a pair of erect, green or slightly purplish 
bracts (spathe), the outer bract rather less than twice the length of the 
inner one, the six spreading segments of the perianth oblong and aristulate 
at the tip. Capsules subglobose, often purplish tinged, about one-fourth 
of an inch in diameter or less. 

In fields and on hillsides, Newfoundland to Saskatchewan, British 
Columbia, Virginia, Nebraska, Colorado and Utah. Flowering from 
May to July. 

There are four other species of blue-eyed grass in New York State: 
S. mucronatum Michaux, 5. arenicola Bicknell, S.grami- 
noides Bicknell and S. atlanticum Bicknell. 


Orchid Family 

Orc hurd a cle ae 
The members of the Orchid family in New York State, of which several 
are described and illustrated here, constitute a very important number 
of our wild flowers and call for a description of the family. They are all 
perennial herbs with corms, bulbs or tuberous roots and entire, sheathing 
leaves, in some species reduced to scales. Flowers perfect, irregular, 
solitary, spiked or racemed. Perianth consisting of six segments, the 
three outer (sepals) similar or nearly so, two of the inner ones (petals) 
lateral, alike; the third inner one (lip) unlike the other two, often markedly 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 26 


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LARGER BLUE FLAG 
Tris versicolor 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 27 


NARROW BLUE FLAG; POISON FLAGROOT 
Iris prismatica 


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WILD FLOWERS OF NEW YORK 65 


so, usually larger, often spurred. Stamens variously united with the style 
into an unsymmetrical column, usually one anther, sometimes two, each 
two-celled; the pollen in two to eight pear-shaped, usually stalked masses 
(pollinia), united by elastic threads, the masses waxy or powdery and 
attached at the base to a viscid disc (gland). Style often terminating in 
a beak (rostellum) at the base of the anther or between its sacs. Stigma 
a viscid surface, facing the lip beneath the rostellum, or the cavity between 
the anther sacs (clinandrium). Ovary inferior, usually long and some- 
times twisted, three-angled, one-celled. Seeds very numerous and minute, 
usually spindle shaped. 
Small White Lady’s-slipper 
Cypripedium candidum Willdenow 
Plate 29 

This is one of the rarer Lady’s-slippers of the east, with rather stiffly 
erect stems 6 to 12 inches high. Leaves three to five, elliptic or lanceolate, 
pointed, 3 to 5 inches long, two-thirds to 14 inches wide; flowers solitary or 
very rarely two ona stem; sepals lanceolate, as long or longer than the lip, 
greenish, spotted with purple; petals somewhat longer and narrower than 
the sepals, wavy-twisted, greenish; lip white, striped with purple or magenta 
inside, about three-fourths of an inch long. 

In marly bogs and low meadows, sometimes in sphagnum bogs, New 
York and New Jersey to Kentucky, Minnesota, Missouri and Nebraska. 
Flowering in June and July. 


Showy Lady’s-slipper 
Cypripedium reginae Walter 
Plate 30 

The largest and most showy of our native orchids, with a stout, villous- 
hirsute stem, I to 3 feet high, leafy to the top. Leaves large, 3 to 8 inches 
long, I to 4 inches wide, elliptic in shape, acute. Flowers 1 to 3; sepals 
round-ovate, white, the lateral ones united for their entire length; petals 
somewhat narrower than the sepals, white, lip much inflated, 1 to 2 inches 
long, white, variegated with crimson and white stripes. 


66 NEW YORK STATE MUSEUM 


In swamps and open wet woods, Newfoundland to Ontario, Minnesota 
and Georgia. Flowering in June and July, more rarely in August in the 
far north. Known also as Whip-poor-will’s Shoe. 

It is doubtful if any wild flower surpasses this in beauty. It has been 
gathered so extensively for its flowers in some localities that it has become 
rather rare. It is one of the plants that above all others needs protection 
in the way of education that will lead lovers of wild flowers to admire its 


beauty where it grows, and to use caution in picking. 


Yellow or Downy Lady’s-slipper 
Cypripedium pubescens Willdenow 
Plate 31 

Stems tall and leafy, 14 to 24 feet high; leaves oval or elliptic 3 to 6 
inches long, 13 to 3 inches wide, pointed; sepals ovate-lanceolate, usually 
longer than the lip, yellowish or greenish yellow and striped with purple, 
petals narrower, usually twisted and elongated; lip much inflated, sub- 
globose, I to 2 inches long, pale yellow to bright yellow and sometimes 
with purple lines, a tuft of white, jointed hairs inside at the top; sterile 
stamen triangular, the thick, somewhat triangular stigma incurved. 

In rich woods and thickets, Nova Scotia to Ontario and Minnesota, 
Alabama and Nebraska. This species is usually regarded as a form of 
Cypripedium parviflorum Salisbury, which has a small, later- 
ally compressed lip one-half to three-fourths of an inch long. Since 
numerous intermediate forms occur it is probable that they represent forms 
of a single variable species. 


Ram’s-head Lady’s-slipper 
Criosanthes arietina (R. Brown) House 
(Cypripedium arietinum R. Brown) 
Plate 32b and Figure V 
Stems 6 to 12 inches high, with three or four elliptic or lanceolate 


leaves 2 to 4 inches long, one-third to 3 inches wide, one-flowered; sepals 


Memoir 15 


WILD FLOWERS OF NEW YORK 


N. Y. State Museum 


Plate 29 


SMALL WHITE LADY’S-SLIPPER 
Cypripedium candidum 


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WILD FLOWERS OF NEW YORK 


(Photograph by G. A. Bailey) 


Figure V 
Ram’s-head Lady’s-slipper 


(Criosanthes arietina (R. Brown) House) 


67 


68 NEW YORK STATE MUSEUM 


separate, lanceolate, one-half to seven-eighths of an inch long, longer than 
the lip; petals linear, greenish brown, about as long as the sepals; lip one- 
half to three-fourths of an inch long, red and white, veiny, prolonged at 
the apex into a long blunt spur, somewhat distorted at the upper end, 
which gives the plant its common name of Ram’s-head Lady’s-slipper. 

A very rare species of cold and damp woods from Quebec to Manitoba, 
Massachusetts, New York and Minnesota. It has been collected but a few 
times in the northern counties of this State. The colored illustration is 
made from a photograph taken in southern Herkimer county by Mr 
Edward H. Eames of Buffalo and the accompanying halftone from a 
photograph by Mr G. A. Bailey of Geneseo. 


Moccasin Flower; Stemless Lady’s-slipper 
Fissipes acaulis (Aiton) Small 
(Cypripedium acaule Aiton) 

Plate 33 and Figure VI 

A short-stemmed plant with only two large, basal, elliptic leaves, 6 to 
8 inches long and 2 to 3 inches wide, thick and dark green. The single 
fragrant flower borne on a scape 6 to 15 inches high; sepals greenish purple, 
spreading, 13 to 2 inches long, lanceolate, the two lateral ones united; 
petals narrower and somewhat longer than the sepals; lip a large, drooping, 
inflated sac with a closed fissure down its whole length in front, 13 to 2} 
inches long, somewhat obovoid, pink with darker veins, rarely white, the 
upper part of the interior surface of the lip crested with long, white hairs. 
Fruit an ascending capsule, pointed at each end. 

In sandy or rocky woods, Newfoundland to Manitoba, south to North 
Carolina, Tennessee and Minnesota. One of the few conspicuous wild 
flowers that appears to be equally at home in the pine lands of the northern 
coastal plain and the rocky woods of the central and northern part of the 
State. On Long Island it sometimes blooms in May but in the north it 
usually blooms in June. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 31 


= 


YELLOW OR DOWNY LADY’S-SLIPPER 
Cypripedium pubescens 


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WILD FLOWERS OF NEW YORK 


(Photograph by E. A. Eames) 


Figure VI 
Moccasin Flower; Stemless Lady’s-slipper 


(Fissipes acaulis (Aiton) Small) 


7O NEW YORK STATE MUSEUM 


(Photograph by O. O. Nylander) 
Figure VII 
Small Round-leaved Orchis 


(Orchis rotundifolia Pursh) 


Small Round-leaved Orchis 
Orchis rotundifolia Pursh 
Figure VII 

Along with the Calypso, this 
small orchis shares the distinction 
of being the rarest wild flower of 
the State. Its slender stem rises 
to a height of 6 to 10 inches and 
bears near the base a single oval 
or orbicular leaf, 1 to 3 inches 
long, with one or two scales 
sheathing the stem below the leaf. 
Flowers in a short terminal spike, 
usually five to ten in number, each 
flower subtended by a small green 
bract. Sepals and petals oval, 
rose-colored, the lateral sepals 
spreading and usually slightly 
shorter, but sometimes longer than 
the petals; lip white, beautifully 
spotted with purple, longer than 
the petals, three-lobed, the middle 
lobe larger, dilated and two-lobed 
or notched at the apex; spur 
slender, shorter than the lip or 
barely equalling it in length. 

Indamp, mossy woods, Green- 
land to the Rocky mountains, 
Maine, New York and Wisconsin. 
The records of its occurrence in 
this State are very few. It has 


been found in Oneida, Herkimer and Lewis counties, but not in recent 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 33 


7 


4 


MOCCASIN FLOWER; STEMLESS LADY’S-SLIPPER 
Fissipes acaulis 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 34 
==. =a 


SHOWY ORCHIS 
Galeorchis spectabilis 


WILD FLOWERS OF NEW YORK Wi 


years. Our illustration is from a photograph by Olaf O. Nylander, taken 
in Maine. 
Showy Orchis 
Galeorchis spectabilis (Linnaeus) Rydberg 
(Orchis spectabilis Linnaeus) 
Plate 34 

Entire plant rather fleshy; stems short, five-angled, 4 to 12 inches 
high from a short rootstock provided with numerous fleshy roots. Leaves 
two, near the base of the stem, obovate, dark glossy green, 4 to 8 inches 
long and 2 to 4 inches wide, clammy to the touch; the three to ten flowers 
in a terminal spike, each flower about 1 inch long, violet-purple mixed with 
lighter purple and white, the subtending bracts sheathing the ovaries; 
sepals united above forming a hood; petals connivent under the sepals and 
more or less attached to them; lip whitish, wavy, produced into an obtuse 
spur, about as long as the petals; column short, scarcely extending above 
the base of the lip, violet on the back. 

In rich woods, New Brunswick to Ontario, Dakota, Georgia, Kentucky, 
Missouri and Nebraska. Flowering in May and June or as late as July in 
the extreme northern part of its range. 


Tall Leafy Green Orchis 
Limnorchis hyperborea (Linnaeus) Rydberg 


Plate 37a 

A rather inconspicuous orchis with small flowers, the stem usually 
stout, 10 inches to 3 feet high. Leaves lanceolate, pointed, 3 to 10 inches 
long, one-half to 12 inches wide. Flowers numerous in a narrow terminal 
spike, small, greenish or greenish yellow; sepals and petals free and spread- 
ing, ovate, blunt; lip entire, lanceolate, blunt, about one-fourth of an 
inch long; spur about as long as the lip, blunt, slightly incurved or often 
thickened at the end (clavate). 

In bogs, wet woods and swamps, Greenland to Alaska, New Jersey, 
Colorado and Oregon. Flowering in May and June or later in cold bogs 
and woods of the north. 


NEW YORK STATE MUSEUM 


“J 
to 


The Tall White Bog Orchis (Limnorchis dilatata (Pursh) 
Rydberg) is usually more slender, with small white flowers; the ovate or 
lanceolate sepals nearly one- fourth of an inch long; petals lanceolate, 
pointed; lip entire, dilated or obtusely three-lobed at the base, blunt at 
the apex, about as long as the blunt and incurved spur. In similar situa- 
tions, but southward only to Maine and New York. 


Large Round-leaved Orchis 
Lysias orbiculata (Pursh) Rydberg 
Plate 35 and Figure VIII 

Flowering scape rather stout, 1 to 2 feet high, with a few inconspicuous 
bracts, and at the base, spreading flat on the ground, two large orbicular 
or orbicular-elliptical, dark-green, shining leaves, silvery beneath, 4 to 8 
inches in diameter. Flowers forming a loose raceme, each flower on a 
pedicel about one-half of an inch long, erect in fruit, greenish white; upper 
sepal short and rounded; lateral sepals spreading, falcate-ovate and blunt; 
one-third to one-half of an inch long; petals smaller and narrower; lip entire, 

oblong-linear, blunt, white, about one-half of an inch long; spur longer 
than the ovary, about 14 to 2 inches long. 

In rich woods, Newfoundland to Minnesota, south to Pennsylvania. 
Flowering in June and July. 

Plate 35 shows the flowers after they have begun to fade. The detail 
of the flowers is shown better in figure VIII. Oakes Ames regards the loose- 
flowered form with long spurs as Habenaria macrophylla 
Goldie. and restricts Lysias orbiculata to the form with dense 
inflorescence and spurs 1 to 1} inches long. 


Hooker’s Orchis 
Lysias hookeriana (A. Gray) Rydberg 
Plate 36 
Stem or flowering scape 8 to 15 inches high, stout, smooth, without 
bracts, but with two fleshy, shining, dark-green, oval-orbicular or obovate, 
spreading or ascending leaves at the base, 3 to 6 inches long, rarely flat on 
the ground. Flowers in a rather loose raceme, 4 to 8 inches long, yellowish 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 


Plate 35 


ir 


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LARGE 


ROUND-LEAVED ORCHIS 
Lysias orbiculata 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 36 


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HOOKER’S ORCHIS 
Lysias hookeriana 


WILD FLOWERS OF NEW YORK 73 


Figure VIII 
Flowers of Large Round-leaved Orchis (Lysias orbiculata (Pursh) 
Rydberg), left; and Large Coralroot (Corallorrhiza maculata 
Rafinesque) at right 


74 NEW YORK STATE MUSEUM 


green; each flower about two-thirds of an inch long; lateral sepals greenish, 
lanceolate and spreading, about one-third of an inch long; petals narrowly 
linear; lip linear-lanceolate, pointed, one-third to nearly one-half of an 
inch long; spur slender, pointed, two-thirds of an inch or more long, as 
long or longer than the ovary. 

In cool, moist woods, Nova Scotia to Minnesota, south to New Jersey, 
Pennsylvania and Iowa. Flowering in June and July or later, in the north. 


Yellow-fringed Orchis 
Blephariglottis ciliaris (Linnaeus) Rydberg 
Plate 37b 

Stem slender, 1 to 23 feet high. Leaves lanceolate, pointed, 4 to 8 
inches long, one-half to 13 inches wide, the upper ones much smaller. 
Flowers orange or yellow, large and showy in a terminal, many-flowered 
spike, 3 to 6 inches long; sepals orbicular or broadly ovate, oblique at the 
base, the lateral ones mostly reflexed; petals much smaller, oblong or 
cuneate, usually toothed; lip oblong, about one-half of an inch long, 
copiously fringed more than halfway to the middle; spur 1 to 14 inches 
long and very slender. 

In meadows and open places in woods, especially in sandy regions, 
Vermont and Ontario to Michigan, Missouri, Florida and Texas. In New 
York State it is rare and local north of the coastal plain, occurring on the 
Schenectady plains and several other places, especially on the Ontario 
lowlands, from Rome west to Lake Erie. 


White-fringed Orchis 
Blephariglottis blephariglottis (Willdenow) Rydberg 
Plate 38 
A plant similar to the Yellow-fringed Orchis, but with a densely or 
rather dense, many-flowered spike of pure-white flowers, the petals toothed 
or somewhat fringed at the apex, rarely entire, the lip copiously or sparingly 


fringed. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 37 


If 7 


B. YELLOW-FRINGED ORCHIS 


A. TALL LEAFY GREEN ORCHIS rae or 
Blephariglottis ciliaris 


Limnorchis hyperborea 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 38 


WHITE-FRINGED ORCHIS 
Blephariglottis blephariglottis 


WILD FLOWERS OF NEW YORK 75 
In bogs and swamps, Newfoundland to Minnesota, Florida and Missis- 
sipp1. When growing with the Yellow-fringed Orchis, it blooms a few days 
earlier. It is found in nearly all the sphagnum bogs of northern New York 
and is not uncommon in moist depressions of the sandy coastal plain. 
Flowering from late June until early August in northern New York. 


The Prairie White-fringed Orchis (Blephariglottis leucophaea 
(Nuttall) Farwell) has larger, white, fragrant flowers, sometimes tinged 
with green; the lip three-parted, the segments broadly wedge-shaped and 
copiously fringed. Most abundant westward but occurring eastward to 
New England. 


Ragged or Green-fringed Orchis 
Blephariglottis lacera (Michaux) Farwell 
Plate 39b 

Stems varying from slender and 1 foot or less high to stout and 2 to 3 
feet high. Leaves firm, lanceolate, 3 to 8 inches long, two-thirds to 12 
inches wide, decreasing in size upward. Flowers greenish yellow or 
greenish, in a loose spike, 2 to several inches long; petals linear, blunt, 
about as long as the sepals; lip three-parted, the segments narrow, deeply 
fringed or lacerate with a few threads about one-half of an inch long; spur 
one-half to three-fourths of an inch long, curved and thickened at the end, 
shorter than the ovary. 

Common in swamps, low meadows and wet woods, Newfoundland to 
Minnesota, south to Georgia and Arkansas. Flowering in June and July. 
Forms frequently occur in which the fringe of the lip is very short. 


Smaller Purple-fringed Orchis 
Blephariglottis psycodes (Linnaeus) Rydberg 
Plate 39a 
Stem rather slender, 1 to 3 feet high. Leaves oval, elliptic or lanceo- 
late, 3 to 10 inches long, two-thirds to 3 inches wide, becoming smaller 
above, dark green and glossy above. Flowers lilac-purple or rarely white, 
fragrant, in a dense, many-flowered terminal raceme, 2 to 8 inches long; 
petals oblong or oblanceolate, toothed on the upper margin; lip three- 


76 NEW YORK STATE MUSEUM 


parted, one-fourth to one-half of an inch broad, the segments fan-shaped 
and copiously fringed, the fringe of the middle segment shorter than that 
of the lateral ones; spur somewhat thickened at the tip, about three-fourths 
of an inch long, and longer than the ovary. 

In meadows, swamps and wet woods, Newfoundland to Minnesota, 
North Carolina and Tennessee. Flowering in July and August. 


The Large or Early Purple-fringed Orchis (Blephariglottis 
grandiflora (Bigelow) Rydberg) has a large, dense raceme of lilac or 
purplish, fragrant flowers, the lower lip, three-parted, the segments broadly 
fan-shaped and copiously fringed to about the middle. The flowers average 
one-third or one-half larger than those of B. psycodes; otherwise they 
are very similar and this may be only a large-flowered race of that species. 
Growing in similar situations. 


Rose Pogonia; Snakemouth 
Pogonia ophioglossoides (Linnaeus) Ker 
Plate 40 

A slender plant with fibrous roots and stems 8 to 15 inches high, 
propagating by runners. Leaves one to three on each stem, one-half to 
4 inches long, lanceolate or ovate, erect, pointed or rather blunt. Flowers 
solitary or sometimes in pairs with leaflike bracts, fragrant, pale rose- 
colored, slightly nodding, 1 to 2 inches broad; sepals and petals similar, 
two-thirds to 1 inch long; lip spatulate, crested and fringed with white. 

In swamps, low meadows and boggy depressions, especially in sandy 
regions, Newfoundland to Ontario, Florida, Kansas and Texas. Flowering 
in June and July, and in the north sometimes in flower as late as August. 


Whorled Pogonia 
Isotria verticillata (Willdenow) Rafinesque 
Figure IX 
Stems 10 to 15 inches high, from long, perennial, horizontal, fleshy 
rootstocks, which give rise to new stems by buds. Stems bearing a whorl 
of five leaves at the summit. Leaves obovate, abruptly pointed at the 


WILD FLOWERS OF NEW YORK Wi 


Figure IX 
Whorled Pogonia 
([sotria verticillata (Willdenow) Rafinesque) 


78 NEW YORK STATE MUSEUM 


apex, sessile, I to 4 inches long, usually only partially developed at flowering 
time. Flowers solitary, erect or declined, on a peduncle one-half to two- 
thirds of an inch long; sepals linear, dull reddish purple, spreading, 14 to 2 
inches long and about one-twelfth of an inch wide; petals linear, erect, blunt, 
light green and arching above the lip, about five-sixths of an inch long; lip 
three-lobed at the end, middle lobe broadest, white and crenulate on the 
margin; two lateral lobes of the lip and the lateral margins tinged and 
veined with bright crimson-purple, most vivid at the apex of the two lateral 
lobes, crest of the lip green and papillose. Capsule erect, 1 to 14 inches 
long. 

In moist soil of woodsand thickets, often around cold sphagnum bogs, 
Ontario and Massachusetts to Michigan, Indiana and Florida. Flowering 
in May and June. 

Arethusa; Dragon’s-mouth; Wild Pink 
Arethusa bulbosa Linnaeus 
Plate 43b 

A low, scapose, smooth plant, 5 to 10 inches high from a small bulb, 
stem bearing one to three loose sheathing bracts on the stem and a single 
linear leaf hidden at first in the upper bract and developing after the flower 
has faded. The single flower arising at the top of the stem from between 
a pair of small unequal scales, rose-purple, 1 to 2 inches high; sepals and 
petals similar, linear to elliptic in shape, obtuse, connivent, hooded, and 
arching over the column; lip usually drooping beneath the sepals and petals, 
the apex broad, variegated with purplish blotches and crested down the 
face with three hairy ridges, the margin fringed or toothed. Capsule about 
t inch long, ellipsoid, strongly six-ribbed. 

In bogs, low meadows and mossy depressions or moist thickets, New- 
foundland to Ontario and Minnesota, South Carolina and Indiana. Flower- 
ing in May and June. One of the most dainty and beautiful of our native 
orchids. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 39 


I = nae a1 


A. SMALLER PURPLE-FRINGED ORCHIS B. RAGGED OR GREEN-FRINGED ORCHIS 
Blephariglottis psycodes Blephariglottis lacera 


Pr er Va a ae 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 40 
— 


ROSE POGONIA; SNAKEMOUTH 
Pogonia ophioglossoides 


WILD FLOWERS OF NEW YORK ie 


Grass Pink; Calopogon 
Limodorum tuberosum Linnaeus 
Plate 41a > 

Scape slender, 12 to 18 inches high from a round, solid bulb, arising 
from the bulb of the previous year, a single leaf appearing the first season, 
followed the next year by the flowering stem and a single linear-lanceolate 
leaf, 8 to 12 inches long, one-fourth to 1 inch wide. Flowers three to fifteen, 
forming a loose, terminal spike or raceme, each about 1 inch broad, purplish 
pink; sepals and petals nearly alike, separate, obliquely ovate-lanceolate, 
acute; lip broadly triangular and dilated at the apex, bearded along the 
face with yellow, orange and rose-colored hairs. 

Common in bogs or boggy meadows. Newfoundland to Ontario and 
Minnesota, south to Florida and Missouri. Flowering in June and July. 


Wide-leaved Ladies’-tresses 
Ibidium plantagineum (Rafinesque) House 
Plate 42a 
A rather inconspicuous little orchid, 4 to 10 inches high with tuberous- 
fleshy roots. Leaves three to five to a stem, mostly near the base, lanceo- 
late, 2 to 5 inches long. Flowers spreading in a dense, terminal spike, 
1 to 2 inches long, one-third to one-half of an inch thick, each flower about 
one-fourth of an inch long; petals and sepals white, the lateral sepals free, 
narrowly lanceolate, the upper sepals somewhat united with the petals; 
lip pale yellow on the face, oblong, not contracted in the middle, the wavy 
apex rounded, crisped or fringed, the base short clawed, bearing mere traces 
of callosities at the base. 
Moist banks, meadows and bogs, Nova Scotia to Minnesota, south to 
Virginia and Wisconsin. Flowers in June and July. 


The most showy species of Ladies’-tresses is Ibidium cernuum 
(Linnaeus) House (figure X), which is 6 to 24 inches high and pubescent 
above. Leaves mainly toward the base of the stem, linear-lanceolate or 
linear, 3 to 14 inches long; flowers very fragrant, white or yellowish, forming 


NEW YORK STATE MUSEUM 


Figure X 


Showy Ladies’-tresses 


(Ibidium cernuum (Linnaeus) House) 


a spike, 4 to 5 inches long 
and one-half to two-thirds of 
an inch thick; flowers spread- 
ing or nodding, each about 
five-twelfths of an inch long in 
three rows; lip oblong or ovate, 
rounded at the apex, crenulate 
or crisped on the margin. 
Flowering in late summer and 
autumn. 

Another common species 
of Ladies’-tresses is Ibidium 
strictum (Rydberg) House, 
perhaps only a race of Ibid- 
ium romanzoffianum, 
which has the sepals and 
petals coherent and connivent 
into a hood. It iscommon in 
bogs and swamps during July 
and August. 


Southern Twayblade 
(Ophrys australis (Lindley) 
House) 


(Listera australis Lindley) 
Figure XI 


The Twayblades are 
among the smallest of our 
native orchids and require 
sharp eyes to detect them 
among the recesses of the 
forest or bogs. The Southern 
Twayblade is about 5 to 10 
inches tall, with two ovate, 
rather pointed, smooth and 
shining sessile leaves slightly 
above the middle of the stem 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 41 
a pine oe se —— 


F > | = —— 


= : — 
A. GRASS PINK; CALOPOGON B. LODDIGES’S RATTLESNAKE PLANTAIN 
Limodorum tuberosum Peramium tesselatum 


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MUOA MAN AO SUAMOTA ATIM 


WILD FLOWERS OF NEW YORK 81 


at the top of which is a loose 
raceme of six to fifteen small, yellow- 
ish green flowers; sepals and petals 
minute, the lip one-fourth to one- 
half of an inch long, and two-parted 
or split nearly to the base four to 
eight times as long as the tiny petals. 

A rare plant of cold, sphagnous 
bogs in the northern counties, west 
to Wayne and Onondaga counties and 
south to Fulton county. In the 
Appalachian region south to Georgia, 
this is not a rare species. 

Very similar to the Southern 
Twayblade is the Heart-leaved 
Twayblade (Ophrys cordata 
Linnaeus) in which the two sessile 
leaves are rounded or slightly heart- 
shaped, and the lip of the flower only 
two or three times as long as the 
petals. It is not rare in the Adiron- 
dack region but on account of its 
small size is easily overlooked. It 
has also been found in deep cedar 
swamps in certain other portions of 
the State. 


Downy Rattlesnake Plantain 
Peramium pubescens (Willdenow) 
MacMillan 


Figure XII 


Flowering scape 6 to 20 inches 


(Photograph by E. A. Eames) 


Figure XI 
Southern Twayblade 


high, densely glandular-pubescent, (O phrys australis (Lindiey) House) 


Figure XII 
Downy Rattlesnake Plantain 


Peramium pubescens (Willdenow) 
(MacMillan) 


| Tennessee. 


NEW YORK STATE MUSEUM 


bearing several lanceolate scales, 
from a branching, fleshy, perennial 
rootstock. Leaves all near the base 
of the scape, I to 3 inches long, two- 
thirds to 1 inch wide, pointed at the 
apex, rather abruptly contracted into 
short petioles, oval or ovate in shape, 
rather strongly reticulated with white 
along the principal veins on the upper 
surface, pale green beneath. Flowers 
white or tinged with green, forming 
a rather dense terminal spike which 
is not one-sided; each flower about 
one-fourth of an inch long or slightly 
less; lateral sepals ovate; upper 
sepals united with the petals to form 


an ovate hood (galea); lip saccate 


with a short broad blunt recurved 


| or spreading tip. 


In dry woods, Maine to Ontario 
and Minnesota, south to Florida and 
Flowering in July and 
August. 


The Lesser Rattlesnake Plantain 
(Peramium ophioides (Fer- 
nald) Rydberg) is only about 6 to 
10 inches high, with smaller, ovate, 
white-blotched leaves and small, 
greenish white flowers, one-eighth 
to one-sixth of an inch long in 
a one-sided spike. Frequent in 
woods. 


WILD FLOWERS OF NEW YORK 83 


Loddiges’s Rattlesnake Plantain 


Peramium tesselatum (Loddiges) 
Heller 


Plate 41b 

Leaves oblong-ovate or ovate- 
lanceolate, I to 2 inches long, one- 
third to one-fourth of an inch wide, 
bright green, abruptly narrowed into 
the petioles which sheathe the base of 
the stem, more or less marked on the 
upper surface with white; stem 6 to 14 
inches high, glandular-pubescent and 
scaly, bearing at the summit a loosely 
spiral spike of flowers I to 5 inches 
long; flowers whitish, lateral sepals 
free, the upper ones united with the 
petals to form a hood (galea), 2 to 3 
lines long, broad and recurved at the 
tip; lip roundish-ovate, slightly saccate 
at the base, the long tip somewhat 
recurved. 

In coniferous woods, more rarely 
in open places, Newfoundland to On- 
tario, Pennsylvania and Michigan. 
Flowering from July to September. 


White Adder’s-mouth 


Malaxis monophylla (Linnaeus) Swartz 
Figure XTII 


St fend LO otinGe Figure XIII 
ber ok eh White Adder’s-mouth 


high, smooth, bearing below the middle (Malaxis monophylla (Linnaeus) 
a single leaf which sheathes the stem, (Swartz) 


84 NEW YORK STATE MUSEUM 


the blade 1 to 3 inches long, one-half to 13 inches wide. Flowers whitish, 
very small, in a slender raceme, 1 to 4 inches long; each flower about one- 
half of an inch long on a very short, nearly erect pedicel; lip triangular or 
ovate and long pointed. 

In woods, thickets and recent clearings, Quebec to Manitoba, south 
to Pennsylvania and Nebraska. Flowering in July. 

The Green Adder’s-mouth (Malaxis unifolia Michaux) has the 
single leaf clasping the stem near the middle; flowers greenish on slender 
pedicels; lp broad and three-toothed at the apex. Small and incon- 
spicuous as well as rather rare. 


Large Twayblade 
Liparis liliifolia (Linnaeus) L. C. Richard 
Figure XIV 

Sceapes 4 to 10 inches high, striate and smooth from a perennial solid 
bulb, the base of the stem sheathed by several scales and two ovate or oval, 
blunt, shining, light-green leaves, 2 to 5 inches long, 1 to 23 inches wide. 
Flowers yellow, numerous, rather showy, forming a loose terminal raceme; 
petals very narrow and threadlike; lip erect, large, fully one-half inch long, 
wedge-obovate in shape. Capsule club-shaped and about one-half inch 
long. 

In moist woods, thickets and recent clearings, Maine to Minnesota, 
south to Georgia and Missouri. Flowering from latter part of May to July. 


Fen Orchis; Loesel’s Twayblade 
Liparts loeselit (Linnaeus) L. C. Richard 
Plate 42b 
A low bog orchis with two light-green, shining, elliptic or elliptic- 
lanceolate leaves, 2 to 6 inches long, one-half to 2 inches wide and obtuse, 
arising with the short stem from a solitary bulb which, with the base of the 
stem, is sheathed by several scales and the remains of leaves of former 


seasons; flowers greenish yellow, about one-fourth of an inch long, in a 


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WILD FLOWERS OF NEW YORK 


Figure XIV 
Large Twayblade 
(Liparis liliifolia (Linnaeus) L. C. Richard) 


85 


86 NEW YORK STATE MUSEUM 


terminal, loosely flowered raceme; petals linear, somewhat reflexed; lip 
obovate, pointed, a little shorter than the petals and sepals, its tip incurved; 
capsules about one-half of an inch long, wing-angled. 

In wet thickets, springy banks, and bogs or boggy meadows, Nova 
Scotia to Saskatchewan, south to Alabama and Missouri. Flowering from 


late May to July and sometimes later. 


Calypso 
Cytherea bulbosa (Linnaeus) House 
(Calypso borealis Salisbury) 
Plate 43a 

Stem or scape 3 to 7 inches high from a perennial bulb one-half of an 
inch or less thick with coralloid roots. The scape bears two or three loose 
sheathing scales and at the base a single round-ovate leaf, 1 to 2 inches 
long, blunt or pointed at the apex and rounded or heart-shaped at the base, 
the petiole 1 to 2 inches long. Flower showy, solitary, 1 to 13 inches broad, 
at the summit of the scape, variegated with purple, pink and yellow; petals 
and sepals similar, nearly equal, linear, erect or spreading, each with three 
longitudinal purple lines. Lip saccate, large, two-divided below, spreading 
or drooping, with a patch of yellow woolly hairs. Column erect, broadly 
ovate and petallike, shorter than the petals, bearing the lidlike anther 
just below the summit. 

In bogs and cold Arbor Vitae swamps, Labrador to Alaska, south to 
Maine, New York, Michigan, California and in the Rocky mountains to 
Arizona and New Mexico. Also in Europe. 

One of the rarest and at the same time most beautiful of our native 
Orchids, appearing much like a small Cypripedium (Lady’s-slipper). In 
New York it has been found in several localities in southern Herkimer, in 
Lewis, Oswego and Onondaga counties. The writer found it several years 
ago in Lodi or Tamarack swamp near Syracuse, but the place has since been 
obliterated by the growth of the city. For the illustration used here we 
are indebted to Edward A. Eames of Buffalo. 


WILD FLOWERS OF NEW YORK 87 


Large Coralroot 
Corallorrhiza maculata Rafinesque 
Figure VIII, page 73 
Stems stout or slender, 8 to 20 inches high, with a large mass of coral- 
like branching rootstocks, the stem purplish, clothed with several appressed 
scales, devoid of green leaves or green color. Flowers ten to thirty, form- 
ing a terminal raceme 2 to 8 inches long, purplish brown; sepals and petals 
linear-lanceolate, about one-fourth of an inch long; lip white, spotted and 
lined with crimson, oval or ovate in outline, deeply three-lobed, crenulate; 
spur yellowish. Fruiting capsules ovoid or oblong, one-half to two-thirds of 
an inch long and drooping. 
In woods, Nova Scotia to British Columbia, south to Florida, Missouri, 
New Mexico and California. Flowering from July to September. 


The Early Coralroot (Corallorrhiza corallorrhiza (Linnaeus) 
Karsten) blooms in May and June. It has smaller flowers of a dull- 
purple color; sepals and petals about one-fourth of an inch long and a 
whitish, oblong lip; spur reduced to a small protuberance adnate to the 
summit of the ovary. Common in moist woods. 

the smallor Vate Coralroot (Corallorrhiza odon tophaze 
(Willdenow) Nuttall) blooms from July to September. It is 6 to 15 inches 
high and very slender. Flowers purplish; sepals and petals about one- 
sixth of an inch long or less, marked with purple lines; lip entire or den- 
ticulate, whitish, spotted with purple. A rather rare plant of moist woods. 
All the species of Coralroot are devoid of green leaves or green coloring 
matter in the stems, because of their parasitic or saprophytic habit. 


Lizard’s-tail Family 
Saururaceae 
Lizard’s-tail 
Saururus cernuus Linnaeus 


Plate 44 
Stem 2 to 5 feet high from a slender rootstock, jointed, pubescent when 
young, becoming smooth. Leaves ovate, thin, palmately five to nine- 
ribbed, dark green, entire, deeply cordate at the base, acuminate, 3 to 6 


88 NEW YORK STATE MUSEUM 


inches long, 2 to 35 inches wide; petioles sheathing the stem at the enlarged 
nodes; the stem terminating in one or two elongated white spikes, their tips 
drooping in flower. Flowers fragrant, small, white, without any perianth; 
stamens six to eight, white, spreading; ovary consisting of three or four 
carpels united at the base, becoming slightly fleshy and strongly wrinkled 
in fruit. 

In swamps and shallow water, Rhode Island to Florida west to southern 
Ontario, Minnesota and Texas. Flowering from June to August. 


Nettle Family 
Urticaceae 
False Nettle 


Boehmeria cylindrica (Linnaeus) Swartz 


Plate 45 

A perennial, rough-pubescent or nearly smooth, erect plant, 1 to 3 
feet tall, not provided with stinging hairs as are the true nettles. Leaves 
mostly opposite, thin, petioled, ovate to ovate-lanceolate, coarsely toothed, 
1 to 4 inches long, one-half to 13 inches wide. Flowers greenish, small and 
dioecious, that is, some of the axillary spikes containing only staminate 
flowers, and others only pistillate flowers, or some of the spikes containing 
both kinds of flowers (androgynous); the staminate spikes interrupted, the 
pistillate mostly continuous, one-fourth to 14 inches long, often terminated 
by small leaves. 

In moist soil and thickets, Quebec and Ontario to Minnesota, Florida, 
Texas and the West Indies. Flowering from July to September. Not an 
attractive plant but figured here as a representative of the Nettle family, 
none of our species having conspicuous flowers. The True or Stinging 
Nettle (Urtica dioica Linnaeus), naturalized in the eastern states 
from Europe, the Slender Wild Nettle (Urtica gracilis Aiton) and 
the Wood Nettle (Urticastrum divaricatum (Linnaeus) Kuntze) 
are all provided with stinging hairs and are most unpleasant plants to 
encounter unexpectedly in the woods. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 44 
= = 


LIZARD’S-TAIL 
SAUruvrus Cernuus 


y 
, 


7 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 45 
Ir RS ee eT a | 


FALSE NETTLE 
Boehmeria cylindrica 


WILD FLOWERS CF NEW YORK 89 


Sandalwood Family 
Santalaceae 
Bastard Toadflax 
Comandra umbellata (Linnaeus) Nuttall 
Plate 28b 

Stems numerous from a horizontal, branching rootstock, said to be 
parasitic on the roots of adjacent herbs, 6 to 18 inches tall, usually branched 
and very leafy. Leaves oblong or oblong-lanceolate, pale green, pointed 
at each end, sessile, one-half to 15 inches long, the lower ones smaller. 
Flowers numerous in terminal cymes, corymbose at the summit of the stem, 
or also axillary, their branches divergent or ascending, greenish white, 
white, or purplish, about one-fifth of an inch long, calyx usually five-lobed, 
corolla none. Fruit a globose drupe about one-fourth of an inch in 
diameter, crowned by the upper part of the calyx tube and its five oblong 
lobes. 

In dry fields and thickets, especially sandy soil, Cape Breton Island to 
Ontario and Assiniboia, south to Georgia, Kansas and Arkansas. At 
Bergen swamp in Genesee county, New York, it grows in an open marl 
bog, which, however, is dry in certain seasons, in company with Arethusa 
bulbosa, Cypripedium candidum, Scirpus caespitosus, 
Anticlea chlorantha, Triantha glutinosa, Solidago hough- 
tonii, and other bog plants, a habitat most unusual for this species. 


Birthwort Family 
NPISEOVO EIA CCA 
Wild or Indian Ginger 
Asarum canadense Linnaeus 
Plate 46 

Leaves clustered in pairs from a slender, branching, aromatic root- 
stock, having the flavor of ginger. Leaf blades long petioled, reniform, 
thin, short pointed at the apex, 3 to 7 inches broad, dark green, the base 


gO NEW YORK STATE MUSEUM 


deeply cordate. Entire plant densely and finely pubescent. Flowers on 
a short, slender peduncle from between the bases of the petioles, often 
concealed or partly buried in old leaves on the ground, I inch or more broad, 
brownish purple; calyx ovoid, its tube completely adnate to the ovary, its 
three lobes inflexed in bud, ovate-lanceolate, acute or long-acuminate, 
spreading; stamens twelve in number, the stout, short filaments inserted 
on the inferior six-celled ovary. 

In rich woods, New Brunswick to Manitoba, south to North Carolina, 
Missouri and Kansas. Flowering in April and May. 

In southern New York occurs also the Short-lobed Wild Ginger 
(Asarum reflexum Bicknell) with smaller flowers, the calyx tube white 
within, and the triangular, acute lobes strongly reflexed. 


Buckwheat Family 
Proll yeecorm a, cierare 


Swamp Smartweed 


Persicaria muhlenbergii (S. Watson) Small 


Plate 47a 


Perennial by long rootstocks, rooting in the mud or in the water, stem 
erect, glabrous, or somewhat pubescent, enlarged at the nodes, 1 to 3 feet 
high. Leaves ovate-lanceolate or oblong-lanceolate, the upper ones often 
narrower, 2 to 8 inches long, acute or acuminate at the apex, rounded or 
cordate at the base, petioled, ocreae cylindric, becoming loose, not ciliate. 
Flowers in linear-oblong, dense, terminal racemes, 1 to 3 inches long; calyx 
dark rose-colored, five-parted; stamens five in number. 

In swamps, marshes and moist soil, Ontario to British Columbia, 
Virginia, Louisiana and Mexico. Flowering from July to September. 


Lady’s-thumb; Heartweed 
Persicaria persicaria (Linnaeus) Small 
Plate 470 
Annual, smooth or somewhat puberulent, stems one-half to 2 feet high, 


erect or nearly so, simple or branched. Leaves lanceolate or linear-lanceo- 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 46 


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WILD OR INDIAN GINGER 
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late, 2 to 6 inches long, nearly sessile, acuminate at both ends, conspicuously 
dotted, usually with a triangular or lunar dark blotch near the center; ocreae 
cylindric, fringed with short bristles. Flowers in dense, erect, ovoid or 
oblong racemes, one-half to 2 inches long, pink to dark purple; stamens 
usually six. 

Native of Europe but naturalized and often an abundant weed in 
waste places throughout North America, except in the extreme north. 


Arrow-leaved Tearthumb 
Tracaulon sagittatum (Linnaeus) Small 
Plate 48a 

Stems slender, weak, annual, decumbent, or climbing over other plants 
by the numerous sharp, recurved prickles which arm its four prominent 
angles. Leaves lanceolate-sagittate or oblong-sagittate, 1 to 3 inches long, 
pointed at the apex, slightly rough margined, the lower leaves petioled, 
upper ones sessile or nearly so, prickly on the petioles and beneath on the 
midribs; ocreae oblique, not ciliate. Flowers in terminal heads or racemes, 
rose-colored or greenish; stamens usually eight in number; style three- 
parted. 

In wet soil, Newfoundland and Nova Scotia to the Northwest Territory, 
south to Florida and Kansas. Flowering from July to September. 


Halberd-leaved Tearthumb 
Tracaulon arifolium (Linnaeus) Rafinesque 
Plate 48b 

Stems angled, reclining, 2 to 6 feet long from a perennial root, armed 
with recurved prickles. Leaves broadly hastate, long petioled, 1 to 8 inches 
long, pubescent or glabrous beneath, the apex and basal lobes sharp pointed ; 
petioles and larger nerves prickly; peduncles and pedicels glandular; ocreae 
oblique, fringed at the summit with short bristles and at the base with 
slender prickles. Flowers in terminal and axillary heads or racemes, rose- 
colored or greenish, four parted. Stamens six; style two-parted. 


g2 NEW YORK STATE MUSEUM 


In moist or wet soil and thickets, New Brunswick and Ontario to 
Minnesota, south to Georgia. Flowering from July to September. 


Climbing False Buckwheat 
Bilderdykia scandens (Linnaeus) Greene 
(Polygonum scandens Linnaeus) 
Plate 15b 

Stems slender or stout, glabrous, high climbing, 2 to 20 feet long from 
a perennial root, somewhat rough on the ridges which mark the stem. 
Leaves ovate, sharp pointed, cordate at the base, 1 to 6 inches long, or the 
upper ones smaller, finely dotted, ocreae oblique, smooth and glabrous. 
Flowers in numerous panicled racemes, 2 to 8 inches long, usually inter- 
rupted with small leaves, yellowish green or whitish; calyx five-parted; 
stamens eight; calyx in fruit about one-half of an inch long with crisped 
wings. 

In woods, thickets and on banks and along fence rows, Nova Scotia 
to Ontario and British Columbia, south to Florida, Nebraska and Texas. 
Flowers in August and September. 


Coast Jointweed 


Polygonella articulata (Linnaeus) Meisner 
Plate 49a 

Stems slender, wiry, erect or somewhat diffusely spreading, annual, 
glaucous, simple or the larger ones often much branched, 4 to 12 inches high. 
Leaves linear or linear-subulate, with revolute margins, sessile, one-third 
to 14 inches long, jointed to the summits of the ocreae. Flowers small, in 
numerous terminal racemes on reflexed pedicels; calyx five-parted, its 
segments white with a conspicuous purple midrib. 

In sand near or on the seashore from Maine to Florida, and inland on 
the Schenectady plains, those east of Oneida lake, and along the Great 
Lakes. 


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WILD FLOWERS OF NEW YORK 93 


Goosefoot Family 
Chenopodiaceae 
Slender or Jointed Glasswort; Saltwort 
Salicornia europaea Linnaeus 
Plate 49b 
A fleshy, glabrous annual plant, 4 to 20 inches high, usually erect and 
much branched, with opposite, ascending branches, their joints two to 
four times as long as thick. Leaves reduced to mere scales. Fruiting 
spikes 1 to 3 inches long. Flowers three at each joint, the middle one as 
high as the lateral ones. Each flower consists of a fleshy, obpyramidal 
three-toothed calyx, two stamens and an ovoid ovary. Seed inclosed by 
the spongy fruiting calyx. 
Common in salt marshes along the coast from Anticosti to Georgia, 
and at the head of Onondaga lake. 


Pokeweed Family 
Phy tola ceaicieare 
Poke; Scoke; Pigeon Berry; Garget 


Phytolacca americana Linnaeus 


Plate 50 

A tall, strong-smelling, succulent and glabrous plant with an erect 
herbaceous stem 3 to 10 feet tall, from a large, perennial, poisonous root, 
the pith of the stem divided into discs separated by lens-shaped cavities. 
Leaves oblong-lanceolate or ovate-lanceolate, pinnately veined, acute or 
acuminate at both ends, petioled, 5 to 12 inches long. Flowers in terminal 
racemes, which become opposite the leaves by continued growth of the 
stem. Each flower about one-fourth of an inch broad, consisting of four 
or five rounded, white sepals; ten stamens, slightly shorter than the sepals; 
and a ten-celled green ovary. Fruit a long raceme of dark-purple berries, 
each one-fourth to three-eighths of an inch in diameter. 

In waste places, fields, woods and thickets, usually in moist soil, often 
in stony fields and frequently a troublesome weed. Distributed from Maine 


94 NEW YORK STATE MUSEUM 


and Ontario to Minnesota, Arkansas, Florida, Mexico and Bermuda. 
Europe has contributed many weeds to America, but the Pokeweed or 
Scoke is one of the few American plants, often reckoned as a weed, which 
has become thoroughly naturalized in many parts of Europe. 


Purslane Family 
Portulacaceae 
Narrow-leaved Spring Beauty 
Claytonia virginica Linnaeus 
Plate sra 


Stems 6 to 12 inches long, ascending or decumbent from a deep, tuber- 
ous, perennial root. Leaves linear or linear-lanceolate, glabrous and some- 
what fleshy, blunt or pointed, narrowed below into a petiole, the lower 
leaves 3 to 7 inches long, one-eighth to one-half of an inch wide, the two- 
stem leaves opposite and shorter. Flowers one-half to seven-eighths of an 
inch broad, white or pink with darker pink veins, in a loose terminal raceme, 
often becoming 3 to 6 inches long. Sepals two, ovate, persistent; petals 
five, emarginate; pedicels slender, becoming recurved. 

In moist woods, Nova Scotia to Saskatchewan, south to Georgia, 
Montana and Texas. Flowering in early spring. More frequently found 
in low woodlands and along streams in the upper part of the State, and not 


so frequent as the next species except near the coast. 


Carolina or Wide-leaved Spring Beauty 


Claytonia caroliniana Michaux 


Plate 69a 

Closely resembling the Narrow-leaved Spring Beauty, but usually 
more erect. Basal leaves ovate-lanceolate or oblong, 1 to 4 inches long, 
one-half to 1 inch wide, obtuse; stem leaves petioled; flowers fewer. The 
corm is usually thicker and depressed at the top. 

In damp woods, Nova Scotia to Saskatchewan, Connecticut, south to 
North Carolina along the mountains, and to Ohio and Missouri. Rare or 
absent near the coast in New York, but very abundant in the interior where 


it occurs in almost every moist woodland. Flowering in the early spring. 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum : Plate 50 


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POKE; SCOKE; PIGEON BERRY; GARGET 
Phytolacca americana 


WILD FLOWERS OF NEW YORK 95 


Chickweed Family 
Alsinaceae 
Field or Meadow Chickweed 
Cerastium arvense Linnaeus 
Plate 63a 

A densely matted or tufted perennial plant, usually more or less 
pubescent. Flowering stems 4 to 10 inches high, simple or sparingly 
branched. Lower leaves and those of the numerous sterile shoots linear- 
oblong, close together, slightly narrowed at the base. Leaves of the 
flowering stems more distant, linear or narrowly lanceolate, one-half to 
13 inches long, one-eighth of an inch wide or slightly wider. Flowers white, 
one-half to three-fourths of an inch broad, several in a cymose inflorescence. 
Petals five, obcordate or deeply notched at the ends, much longer than the 
lanceolate, acute sepals. Fruit a small, cylindrical capsule, a little longer 
than the sepals and slightly oblique at its apex. 

In dry, rocky places, stony fields or ledges, Labrador to Alaska, south 
to Georgia, Missouri, Nevada and California. Also found in Europe and 
Asia. Flowering from April to July. In New York usually in bloom 
during some part of May. 

This is a much larger flowered and more attractive plant than the 
Mouse-ear Chickweed (Cerastium vulgatum Linnaeus), which is 
naturalized almost everywhere, especially in the stony or rocky places 


where the Field Chickweed is found. 


Pink Family 
Cra riy.onpiieyolelaxcierane 
Bladder Campion; White Ben 
Silene latifolia (Miller) Britten & Rendel 


Plate 52 
Stems herbaceous, from a perennial root, glaucous and glabrous, or 
rarely pubescent, extensively branching from the base, 6 to 20 inches high. 


96 NEW YORK STATE MUSEUM 


Leaves opposite, ovate-lanceolate, pointed, variable in size, the lower ones 
often spatulate. Flowers white, two-thirds to seven-eighths of an inch 
broad, in loose cymose panicles, often drooping; calyx tubular-campanulate, 
becoming globose and much inflated, about one-half of an inch long, strongly 
veined, with five triangular, acute lobes; petals five, each two-cleft, with or 
without a small crown. 

In fields, meadows and waste places, New Hampshire to Ontario, 
New Jersey and Missouri. Native of Europe and naturalized in this 
country. Flowering in July and August. 


Wild Pink 


Silene caroliniana Walter 


Plate 53 

Densely tufted and perennial from a stout root, 4 to to inches high, 
viscid-pubescent, especially above, often glabrous below. Basal and lower 
leaves spatulate or oblanceolate, pointed or blunt, 2 to 4 inches long; stem 
eaves sessile, shorter, oblong or lanceolate. Flowers pink, about 1 inch 
broad, in terminal cymes. Calyx narrow and tubular, much enlarged by 
the ripening pod, its teeth ovate, pointed; petals cuneate, emarginate, eroded 
or finely toothed at the apex, crowned at the base of the claw. 

In dry, sandy or rocky soil, Maine to Georgia, west to central New 


York, Pennsylvania and Kentucky. 


Cuckoo-flower; Ragged Robin 
Lychnis flos-cuculi Linnaeus 
Plate 54a 
Stems slender, erect, 1 to 2 feet high, simple or branching, from a 
thick, perennial root, downy-pubescent below, slightly viscid above. Lower 
and basal leaves spatulate or oblanceolate, 2 to 4 inches long; upper leaves 
lanceolate or linear-lanceolate, opposite and sessile, the uppermost leaves 
reduced to small bracts; flowers pink, purplish, blue, or white, three- 
fourths to 1 inch broad, in many-flowered panicles at the summit of the 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 51 


r 


A. NARROW-LEAVED SPRING BEAUTY 
Claytonia virginica 


> 


B. ROUND-LOBED HEPATICA OR LIVERLEAF 
Hepatica hepatica 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 52 
= 


BLADDER CAMPION; WHITE BEN 
Silene latifolia 


WILD FLOWERS OF 


Memoir 15 N. Y. State Museum 


NEW YORK 


Plate 53 


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WILD PINK 


Silene caroliniana 


WILD FLOWERS OF NEW YORK 97 


stems; calyx one-fourth of an inch long, ten-nerved, cylindrical, becoming 
campanulate in fruit, its apex with five triangular, pointed teeth; petals 
five, narrowly clawed, the spreading limbs each cleft into four linear lobes, 
of which the middle pair is longest. 

Moist meadows, fields and waste places, New Brunswick to New 
Jersey and Pennsylvania. Frequent in cultivation. A native of Europe 
but thoroughly naturalized in many places in the eastern states. Flowering 
in July and August. 

Sacred Bean Family 
Nelumbonaceae 
American Nelumbo or Lotus 
Nelumbo lutea (Willdenow) Persoon 
Plate 55 

Rootstock stout, nearly horizontal, tuberiferous, in mud beneath 2 to 
6 feet of water. Leaves 1 to 2 feet broad, orbicular, or somewhat con- 
stricted in the middle, centrally peltate, floating or raised a foot or two 
out of the water, prominently veined, smooth and dark green above, more 
or less pubescent and finely scaly beneath; leaf petioles and flower stems 
3 to 7 feet long, rigid and tough, with several large air canals. Flowers 
pale yellow, fragrant, 4 to 10 inches broad; petals concave, obovate, blunt, 
numerous, surrounded by four or five overlapping scales. Carpels 
numerous, contained in pits in the large convex receptacle which becomes 
3 to 4 inches long and obconic in fruit. Seeds about one-half of an inch 
in diameter. 

In rivers and lakes, locally distributed from Massachusetts to Minne- 
sota, Nebraska, Louisiana and Cuba. Flowering in August at Sodus bay, 
where a large colony of it exists. Tubers and seeds farinaceous and edible. 
The local distribution in the north may be due to introduction by the 
Indians. The Indian Lotus or Sacred Bean (Nelumbo nelumbo 
(Linnaeus) Karsten), with large pink flowers, is frequent in cultivation. 


98 NEW YORK STATE MUSEUM 


Water Lily Family 
Nymphaeaceae 
Large Yellow Pond Lily; Spatter-dock 
Nymphaea advena Solander 
Plate 56 

Floating and emersed leaves 5 to 12 inches long and 5 to g inches 
broad, ovate or orbicular-oval, thick, with a sinus 2 to 5 inches deep and 
generally open; submerged leaves, when present, thin and membranous; 
petioles, peduncles and lower surfaces of the leaves usually pubescent. 


Flowers 14 to 23 inches broad, depressed, globose, yellow, usually tinged 


with purple within; sepals six, oblong; petals fleshy, oblong truncate, one- 
half to two-thirds of an inch long; stamens numerous in five to seven rows; 
carpels numerous, united into a compound pistil which is surmounted by 
an undulate, yellow or pale-red stigmatic disc with twelve to twenty-four 
rays, ripening into an ovoid, berrylike fruit, 1 to 2 inches long and about 
I inch thick, maintained at the surface of the water or above it. 

In ponds, lakes, slow streams or often subterrestrial in boggy meadows, 
Labrador and Nova Scotia to the Rocky mountains, south to Florida, Texas 
and Utah. Flowering from May to September. Consists of several races 
or perhaps species, differing in the character of the pistil, stigmatic disc 
and leaf outline. In the lakes and ponds throughout the north the small 
Yellow Pond Lily (Nymphaea microphylla Persoon) is also 
found, with flowers 1 inch broad or less, and small leaves 2 to 4 inches long 
and 1 to 3 inches broad. The northern form of the larger Yellow Pond 
Lily 1s described in some books under the name cf Nymphaea vari- 
egata (Morong) Greene, and a hybrid between the two, Nymphaea 
rubrodisca (Morong) Greene, is of frequent occurrence, having fewer 
stigmatic rays than N. variegata and spatulate petals. Our illustra- 
tion is from a plant on Long Island. 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 55 


AMERICAN NELUMBO OR LOTUS 
Nelumbo lutea 


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WILD FLOWERS OF NEW YORK 99 


Sweet-scented White Water Lily 
Castalia odorata (Aiton) Woodville & Wood 
Plate 57 

Aquatic, with a thick, horizontal rootstock. Leaves floating, orbicular 
or nearly so, 4 to 12 inches in diameter, glabrous, green and shining above, 
purple and somewhat pubescent beneath, the sinus open or almost closed, 
petioles and peduncles slender with four main air channels. Flowers white 
or in some varieties pink or rose-colored, 3 to 6 inches broad, very fragrant, 
with four greenish sepals and numerous, narrowly oblong, blunt petals, the 
inner ones shorter and narrower and gradually passing into stamens; 
stamens numerous and yellow. Carpels numerous, united into a compound 
pistil with radiating, linear, projecting stigmas, becoming a globose, fleshy 
fruit and ripening beneath the surface of the water by the coiling of the 
peduncles. 

In ponds, lakes and slow streams, Newfoundland to Manitoba, south 
to Florida, Louisiana and Kansas. Flowering from June to August. 


Tuberous White Water Lily 
Castalia tuberosa (Paine) Greene 
Plate 58 

Leaves orbicular, 5 to 12 inches in diameter, floating, green on both 
sides, sometimes slightly pubescent beneath, sinus open or closed. Root- 
stock thick, with numerous lateral tuberous-thickened branches which 
become detached and propagate the plant. Flowers white, 4 to 9 inches 
broad, slightly fragrant or inodorous; petals oblong, broader than those of 
C. odorata, obtuse. Fruit depressed-globose. 

In shallow water of bays and protected coves, Lake Champlain, Lake 
George, Oneida lake and along the Great Lakes to Michigan, south to 
Delaware and eastern Nebraska and Arkansas. Flowering in July and 
August. 


100 NEW YORK STATE MUSEUM 


Crowfoot Family 
Ranunculaceae 
Marsh Marigold; Cowslip 
Caltha palustris Linnaeus 
Plate 59 

A succulent, herbaceous plant with stout, glabrous, hollow stems, 
erect or ascending, 1 to 2 feet high, branching and bearing several or 
numerous bright-yellow flowers. Lower leaves long petioled, the blades 
cordate or reniform, 2 to 8 inches broad, with a narrow sinus, crenate, 
dentate, or nearly entire on the margin. Upper leaves smaller, with short 
petioles or sessile with nearly truncate bases. Flowers 1 to 13 inches 
broad; sepals oval, obtuse, petallike. True petals none. Stamens numer- 
ous, obovoid. Carpels several, in fruit forming follicles which are 
slightly compressed, about one-half of an inch long, and slightly curved 
outward. 

In swamps, wet meadows and marshes, Newfoundland to South 
Carolina, west to Saskatchewan and Nebraska. Flowering in May and 
June. 

Goldenseal; Orange-root 
Hydrastis canadensis Linnaeus 
Figure XV 

An erect, perennial herb with a thick yellow rootstock. Stems 10 to 
15 inches high. Usually each plant with a single, long-petioled basal leaf 
which is 5 to 8 inches broad and palmately 5 to 9-lobed, the lobes broad, 
pointed, sharply and unequally toothed. Stem leaves two, borne at the 
summit of the stem, similar in shape but smaller than the basal leaf, the 
uppermost leaf just below the solitary, greenish white flower, which is 
one-third to one-half of an inch broad. Sepals three, petallike and falling 
away as the flower opens. Petals none. Stamens numerous, their filaments 
widened and about one-sixth of an inch long; anthers oblong, obtuse. 


Carpels several, ripening into an ovoid, crimson head of fruit about two- 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum : 7 Plate 57 


SWEET-SCENTED WHITE WATER LILY 
Castalia odorata 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 58 


| | 


TUBEROUS WHITE WATER LILY 
Castalia tuberosa 


WILD FLOWERS OF NEW YORK IOI 


thirds of an inch long, each carpel in fruit tipped with a short recurved 
beak. 
In moist or low woodlands, Connecticut to Minnesota, Ontario, Kansas 


and Missouri, south to Georgia. 


Figure XV 
Goldenseal or Orange-root 
(Hydrastis canadensis Linnaeus) 


American Globeflower 
Trollius laxus Salisbury 


Plate 61 
Stems few or several from a perennial fibrous root, erect or ascending, 


often slightly fleshy but weak, from a few inches to 2 feet long. Lower 


102 NEW YORK STATE MUSEUM 


leaves long-stalked; upper ones short-stalked or sessile; blades 3 to 5 inches 
broad, parted into five to seven wedge-shaped, cleft or toothed segments, 
the blades small at flowering time, much enlarged later. Flowers 1 to 14 
inches broad, with five to seven spreading, yellowish green, petallike sepals; 
true petals minute, fifteen to twenty-five in number and much shorter than 
the numerous yellow stamens. Fruit about 1 inch broad, consisting of 
several small pods (follicles) each about one-fourth of an inch long and 
tipped with a straight, slender beak of about one-fourth its length. 

A rare or local plant of low or swampy woodlands, New England to 
Delaware, central and western New York to Michigan. Flowering from 
April to June. 

Goldthread 
Coptis trifolia (Linnaeus) Salisbury 
Plate 60a 

A low, herbaceous plant with a slender or filiform bright-yellow, bitter 
rootstock. Leaves all basal, evergreen, long petioled, the blade reniform, 
I to 2 inches broad, divided to the petiole into three wedge-shaped, obtuse 
segments, dark green, shining above, paler beneath, sharply toothed. 
Scape one-flowered, slender; sepals five to seven, oblong, obtuse, white; 
petals small and club-shaped; carpels three to seven, spreading, about 
one-fourth of an inch long, on stalks of about their own length, tipped 
with a beak. 

In damp, mossy woods and bogs, Newfoundland to Virginia and 
eastern Tennessee, Iowa, Minnesota and Alaska. 

In the Memoirs of Bastram and Marshall, page 20, it is stated that 
John Ellis, the eminent naturalist, in a letter to Linnaeus, dated London, 
April 25, 1758, says: “‘ Mr Colden of New York, has sent Dr Fothergill 
a new plant, described by his daughter (Miss Jane Colden). It is called 
Fibraurea, gold thread. This young lady merits your esteem and does 
honor to your system. She has drawn and described 400 plants in your 
method only. She uses the English terms. Her father has a plant called 
after him, Coldenia; suppose you should call this Coldenella, or any other 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 59 


MARSH MARIGOLD; COWSLIP 
Caltha palustris 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 60 
; =a 


A. GOLDTHREAD 
Coptis trifolia 


B. HISPID BUTTERCUP 
Ranunculus hispidus 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 61 


AMERICAN GLOBEFLOWER 
Trollius laxus 


- 


= 7 a 


Foe ee ee Ce” eS ee ee 


WILD FLOWERS OF NEW YORK 103 


name that might distinguish her among your genera.’’ Linnaeus, however, 
referred the plant to his genus Helleborus, and when it was subsequently 
ascertained to be distinct, Salisbury, regardless alike of gallantry and 
justice, imposed on it the name of Coptis. 


Red Baneberry; Black Cohosh 


Actaea rubra (Aiton) Willdenow 
Plate 62 


Stems erect, I to 2 feet high, from a perennial root, pubescent or 
smooth. Leaves ternately divided, the divisions pinnate with the lower 
ultimate leaflets sometimes again compound; leaflets ovate or the terminal 
ones obovate, toothed or more or less cleft or incised with pointed or 
rounded teeth. Flowers small in a dense terminal, ovoid raceme; sepals 
three to five, petaloid and fugacious. Petals four to ten, spatulate, shorter 
than the numerous white stamens; pedicels slender, one-half to two-thirds 
of an inch long. Fruit consisting of a raceme of bright-red, oval or 
ellipsoid berries, each berry about one-half of an inch long. 

In woods, thickets and shaded banks, Nova Scotia to New Jersey and 
Pennsylvania, west to South Dakota and Nebraska. Flowering from April 
to early June. A variety with red berries on slender pedicles (Actaea 
neglecta Gillman) is occasionally found. 


White Baneberry; Snakeroot 
Actaea alba (Linnaeus) Miller 
Plate 63b 
Resembling the Red Baneberry in general habit and aspect. Leaflets 
usually more cut and the teeth and lobes sharply pointed. Flowers in 
oblong racemes; petals truncate at the apex; fruiting pedicels as thick as 
the peduncle or in fruit even thicker, with swollen ends, often reddish; 
berries short-oval, white, sometimes purplish at the ends. A variety with 
berries on thickened pedicels is occasionally seen. 
In rich woods, Nova Scotia to Georgia, west to Minnesota and Mis- 
souri. Flowering in April and May or as late as the middle of June. 


104 NEW YORK STATE MUSEUM 


Biack Snakeroot; Black Cohosh 


Cimicifuga racemosa (Linnaeus) Nuttall 


Plate 64 

Stems tall and slender, 3 to 8 feet high, leafy above, rootstock thick 
and perennial. Leaves ternate with pinnate divisions; leaflets ovate or 
oblong, the terminal ones usually obovate, pointed at the apex, truncate 
or narrowed at the base, margins coarsely toothed, cleft or divided, rather 
thick texture, smooth or nearly so. Flowers white, with a somewhat 
fetid odor, in tall, terminal, simple or compound racemes, 6 to 30 inches 
long, each flower about one-half of an inch broad; petals four to eight, 
two-cleft; stamens very numerous; pistils one or two, sessile. Fruiting 
follicles oval, about one-fourth of an inch long, minutely beaked. 

In woods and shaded rocky places, Maine and Ontario to Wisconsin, 
south to Georgia and Missouri. Flowering in July and August. 


Wild Columbine; Rock Bells 


Aquilegia canadensis Linnaeus 
Plate 65 
Erect and branching, glabrous or somewhat pubescent, 1 to 2 feet 
high, from a perennial root. Leaves ternately decompound, the lower and 
basal leaves slender-petioled, 4 to 8 inches broad, the ultimate leaflets 
I to 2 inches broad, sessile or short stalked, obovate, obtuse, cuneate, 
obtusely lobed and toothed, pale beneath; leaves of the upper part of the 
stem lobed or divided. Flowers nodding, 1 to 2 inches long, scarlet or 
rarely white or yellow; sepals five, regular, petaloid; petals concave, 
produced backward between the sepals into hollow, nearly straight spurs, 
one-half to three-fourths of an inch long, thickened at the end; stamens 
numerous, with the styles long exserted. Fruit erect, consisting of five 
united carpels with slightly spreading filiform beaks. 
In rocky woodlands and clearings, Nova Scotia to the Northwest 
Territory, south to Florida and Texas. Flowering from the latter part of 
April until June. 


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WILD FLOWERS OF NEW YORK 105 


Known in many localities as Honeysuckle, a name which should more 
properly be applied to species of the genus Lonicera. The European 
Columbine (Aquilegia vulgaris Linnaeus) with showy, blue, purple 
or white flowers is an occasional escape from gardens into woods and fields. 


Tall Anemone; Thimbleweed 


Anemone virginiana Linnaeus 
Plate 66 


Stem stout, hairy, 2 to 3 feet tall, branching above at the involucre 
which is composed of two to five, usually three, short-petioled leaves; the 
lateral peduncles often bearing secondary involucres of smaller leaves. 
Basal leaves long petioled, broader than long, three-parted, the divisions 
broadly cuneate-oblong, variously cleft and divided into acute, serrate 
lobes; those of the involucres similar. Flowers two-thirds to 14 inches 
broad, on peduncles 4 to 10 inches long; sepals five, white, obtuse; stamens 
numerous, shorter than the sepals. Carpels numerous, forming an oblong 
to subceylindric head, two-thirds to 1 inch long and about one-half as thick. 

In woods, thickets and clearings, Nova Scotia to Alberta, south to 
South Carolina, Arkansas and Kansas. Flowering from May to July or 
August. Consists of several races, differing in size and color of flower, shape 
of fruit, and in the styles. Of these, the most distinct is Anemone 
riparia Fernald, with smaller greenish flowers and pointed sepals. 

The Slender-fruited Anemone (Anemone cylindrica A. Gray), 
which is frequent in sandy woods and thickets in the eastern states has 
silky hairy stems and tufted basal leaves with narrower divisions, white or 
greenish flowers about three-fourths of an inch broad and the head of fruit 


cylindric, 1 inch long or often longer and one-fourth to one-third as thick. 


Canada or Round-leaved Anemone 


Anemone canadensis Linnaeus 
Plate 67 


Stems slender or stout, 1 to 2 feet tall, somewhat hairy, especially on 
the lower surfaces of the leaves, branching at the involucre. Basal leaves 


106 NEW YORK STATE MUSEUM 


long petioled, broader than long, three to five-pointed, the divisions broad, 
oblong, acute, variously cleft or toothed, those of the primary and secondary 
involucres similar but sessile. Flowers 1 to 13 inches broad; sepals white 
or sometimes tinged with pink, oblong, obtuse; head of fruit globose, con- 
sisting of numerous flattened, orbicular, pubescent achenes, tipped with 
stout, persistent styles. 

Low grounds, along roadsides, railroads and in open woods, Labrador 
to Assiniboia, Massachusetts, Maryland, Illinois, Kansas and Colorado 
Very closely related to Anemone dichotoma of Siberia. Flower- 
ing from May to August, by the development of secondary involucres. 


Windflower; Wood Anemone 


Anemone quinquefolia Linnaeus 
Plate 68a 
Smallest of our wild anemones. Stems simple, glabrous, 4 to 10 inches 


high, from horizontal, perennial rootstocks. Basal leaves long petioled, 
usually developing after the flowering stem, five-parted, the divisions 
oblong, cuneate, dentate; those of the single involucre on slender petioles 
one-half to 1 inch long, three- to five-parted, the divisions about 1} inches 
long, acute, variously cut and lobed. Flowers solitary, about 1 inch broad; 
sepals four to nine, obovate or oval, white or purplish without; head of 
fruit globose, inclined, consisting of several pubescent, oblong achenes, 
tipped with hooked styles. 

Common in moist or low woodlands, Nova Scotia to Minnesota, south 
to Georgia and Tennessee. Flowering in April and May. 


Round-lobed Hepatica or Liverleaf 
Hepatica hepatica (Linnaeus) Karsten 


Plate 51b 
Leaves long petioled, arising with the flowering scapes directly from 
the fibrous roots, reniform, hairy, 2 to 23 inches broad when mature, 
spreading on the ground, three-lobed (occasionaily the lateral divisions 
again lobed), obtuse. Flowers blue, purple or white, one-half to 1 inch 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 64 


BLACK SNAKEROOT; BLACK COHOSH 
Cimicifuga racemosa 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 65 
Te 1 


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WILD COLUMBINE; ROCK BELLS 
Aquilegia canadensis 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 66 


7 


TALL ANEMONE; THIMBLEWEED 
Anemone virginiana 


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WILD FLOWERS OF NEW YORK 107 


broad on hairy scapes 4 to 6 inches high. Each flower subtended by an 
involucre of three sessile, obtuse, oblong, small leaves immediately under 
the flowers. Sepals oval or oblong, obtuse, longer than the numerous 
stamens. Fruit consisting of several oblong, acute, hairy achenes. 

In woods, often in large tufts. Nova Scotia to northern Florida, west 
to Manitoba, Iowa and Missouri. Also in Alaska, Europe and Asia. Flow- 
ering in earliest spring, with us usually early in April but sometimes in 
March, and even unseasonably warm spells in midwinter may find it in 
flower. The leaves of Hepatica hepatica in central Europe 
possess blunt lobes, while our form usually has perfectly rounded lobes. 


Sharp-lobed Hepatica or Liverleaf 
Hepatica acutiloba DeCandolle 


Plate 60b 

Resembling in most respects the Round-lobed Hepatica but the leaf 
lobes and the leaves of the involucre are pointed, the scapes usually a little 
longer, and the flowers somewhat larger. The flowers are often dioecious. 

In woods, Maine, Quebec and Ontario; south in the Alleghanies to 
Georgia; west to Missouri and Minnesota. Rare or absent near the 
Atlantic coast. 

Puzzling forms sometimes occur which are intermediate between this 


and the preceding species. 
Rue Anemone 


Syndesmon thalictroides (Linnaeus) Hoffmannsegg 
Plate 68b 


Stems slender and weak, glabrous, 4 to 10 inches high; the flowering 
stem appearing in early spring from a cluster of tuberous roots, the 
ternately compound basal leaves appearing later and resembling those 
of the Meadow Rue, but smaller. Leaves of the involucre similar, sessile, 
the leaflets long petioled. Flowers perfect, few or several forming a loose 
umbel immediately above the involucre, white or pinkish, one-half to 1 
inch broad; sepals five to ten, thin and soon falling, longer than the 


numerous stamens. 


108 NEW YORK STATE MUSEUM 


In woods, New Hampshire and Massachusetts to Florida, Ontario, 


Minnesota and Kansas. Flowering in early spring, March to early June. 


Swamp or Marsh Buttercup 
Ranunculus septentrionalis (Linnaeus) Poiret 
Plate 70 

Stems branching, 1 to 2 feet long, or becoming longer in summer, 
ascending, the later branches procumbent and often rooting at the nodes. 
Roots simply fibrous; stems glabrous or pubescent. Leaves large, petioled, 
three-divided, the divisions mostly stalked, usually cuneate at the base, 
cleft into broad lobes; petioles of the lower leaves sometimes a foot long. 
Flowers bright yellow, 1 to 14 inches broad; petals five, obovate, twice as 
long as the spreading sepals. Stamens numerous. Fruit a globose or oval 
head of flat, strongly margined achenes, each achene tipped by the subulate, 
persistent, sword-shaped style. 

Marshes, swamps, ditches and low meadows, New Brunswick to Mani- 
toba, Georgia and Kansas. Flowering from April to July. 


Hispid Buttercup 
Ranunculus hispidus Michaux 
Plate 60b 

Plant usually hairy when young, sometimes merely appressed- 
pubescent or glabrate when old; stems ascending or spreading, usually 
several from a thickened, fibrous, perennial root; at flowering time the 
stems only a few inches long, later becoming 1 to 2 feet long, but not 
stoloniferous. Leaves pinnately three to five divided, the divisions 
ovate, oblong or obovate, narrowed or cuneate at the base, sharply cleft 
or lobed, usually thin; flowers one-half to 13 inches broad; petals usually 
five, oblong, about twice as long as the spreading sepals and entire or 
sometimes slightly notched at the apex; achenes of fruit oval, lenticular, 
narrowly margined, abruptly tipped by a subulate style of about one-half 
their length. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 


Plate 67 


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CANADA OR ROUND-LEAVED ANEMONE 


Anemone canadensis 


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WILD FLOWERS OF NEW YORK 109 


In dry woods and thickets, Vermont and Ontario to North Dakota, 
south to Georgia and Arkansas. Flowering from March to May. 


Stiff White Water Crowfoot 
Batrachium circinatum (Sibthorp) Reichenbach 
Plate 79a 

Plant entirely submerged, except the flowers. Stems branching, 
usually 1 foot long or longer. Leaves about 1 inch long, spreading nearly 
at right angles from the stem, only slightly or not at all collapsing when 
drawn from the water, repeatedly forked into capillary divisions. Flowers 
white, one-third of an inch broad on stout peduncles, 1 to 2 inches long 
opposite the leaves, flowering just above the surface of the water; sepals and 
petals five; petals oblong-oval and blunt. Fruit a small cluster of tiny, 
apiculate achenes. 

In ponds and slow streams, Nova Scotia to British Columbia, south 
to North Carolina, also in Europe and Asia. Flowering from June to 
September. 

The form illustrated here has the beak of the achenes about 1 mm long, 
so that it should be classed as Batrachium longirostre (Godron) 
F. Schultz. 

Early Meadow Rue 
Thalictrum dioicum Linnaeus 
Plate 71 

Stems glabrous, erect, 1 to 2 feet high, slender and leafy from brown 
perennial roots. Leaves three to four-ternate. Leaflets thin, pale beneath, 
orbicular or broader, often cordate and the terminal one somewhat cuneate 
five to nine-lobed. Flowers dioecious, greenish or greenish yellow, drooping 
or spreading; panicle elongated, of numerous lateral corymbs or umbels; 
sepals usually four, blunt; petals none; stamens numerous, filaments longer 
than the sepals; anthers linear, blunt, longer than the filaments. Achenes 
in fruit ovoid, sessile or minutely stipitate, strongly ribbed, much longer 
than the style. 


1 0) NEW YORK STATE MUSEUM 


In woods and on shaded banks, Maine to Alabama, Saskatchewan and 
Missouri. Flowering in April and May. 


Fall Meadow Rue 


Thalictrum polygamum Muhlenberg 


Plate 72 

Stems stout, smooth or pubescent but not glandular or waxy, 3 to Io 
feet high, branching. Leaves three to four-ternate; leaflets thickish, light 
green above and pale beneath, oblong or orbicular with three main apical 
pointed or blunt lobes; panicle compound, leafy, a foot long or more. 
Flowers polygamous, white or purplish, usually the pistillate flowers pur- 
plish and the staminate flowers white; filaments broad, narrowly clavate; 
anthers oblong, short. Fruiting achenes ovoid, sessile or short-stipulate, 
six to eight-winged, glabrous or pubescent. 

Marshes, open sunny swamps and low meadows. Newfoundland to 
Florida, Ontario and Ohio. Flowering from July to September. 


Virgin’s Bower; Woodbine; Wild Clematis 
Clematis virginiana Linnaeus 
Figure XVI and Plate 73 

A long vine, climbing over bushes in low woodlands, and along fences 
and watercourses. Stems somewhat woody below but only the root peren- 
nial in the north at least. Leaves opposite, glabrous, trifoliate; leaflets 
broadly ovate, acute at the apex, toothed or lobed, sometimes slightly 
cordate. Flowers in leafy panicles, white, polygamo-dioecious, two- 
thirds to 1} inches broad when expanded. Sepals usually four, spreading, 
petallike; petals none; stamens numerous, spreading; filaments glabrous; 
pistils numerous. In fruit the styles become an inch long or more, 

plumose and persistent on the achenes (figure XVI). 
Nova Scotia to Georgia, west to Manitoba and Tennessee. Flowering 


in midsummer, July to September. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 70 


y 


SWAMP OR MARSH BUTTERCUP 
Ranunculus septentrionalis 


WILD FLOWERS OF NEW YORK 


Figure XVI 
Virgin’s Bower (Clematis virginiana Linnaeus), in fruit 


130 


I12 NEW YORK STATE MUSEUM 


Purple Virgin’s Bower 
Atragene americana Sims 
(Clematis verticillaris De Candolle) 
Figure XVII 
A trailing or partly climbing vine, somewhat woody and perennial 
below, glabrous or nearly so. Leaves trifoliate; leaflets thin, ovate, acute, 
toothed or entire and more or less cordate; petioles and petiolules slender. 
Flowers purplish blue, 2 to 4 inches broad, solitary on slender peduncles 
in the axils of the leaves or at the ends of the branches. Sepals four, thin 
and translucent, strongly veined, silky along the margins and veins; petals 
four, spatulate, one-half to two-thirds of an inch long; stamens very 
numerous, the outer ones usually with broadened filaments; styles long, 
persistent, plumose throughout and about 2 inches long in fruit. 
Rocky woodlands and thickets, Hudson bay to Manitoba, south to 
Connecticut, Virginia and Minnesota. Flowering in May and June. One 
of our rarest wild flowers. 


Erect Silky Leather Flower 


Viorna ochroleuca (Aiton) Small 


Plate 74 

Stems erect, silky-hairy, 1 to 2 feet high and somewhat woody at the 
base from a thickened, woody, perennial root. Leaves opposite, simple, 
sessile, ovate or elliptical-ovate, blunt, smooth and glabrous above, silky 
and reticulate-veined beneath, entire or rarely somewhat lobed; each stem 
with a single terminal nodding flower about 1 inch long or less. Calyx 
rather broadly cylindric in shape, composed of four or five thick sepals, 
very silky without, their yellowish-green tips recurved; petals none. 
Stamens numerous, parallel with the sepals, their anthers very narrow. 
Pistils very numerous, their styles silky or plumose. In fruit the fleshy 
sepals fall away leaving an erect head of small. achenes plumose with the 


long, yellowish-brown, persistent styles which are 1 to 2 inches long. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 71 


EARLY MEADOW RUE 
Thalictrum dioicum 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Ss Plate 72 S 


FALL MEADOW RUE 
Thalictrum polygamum 


WILD FLOWERS OF NEW YORK I13 


Figure XVII 
Purple Virgin’s Bower 


(Atragene americana Sims) 


Ii4 NEW YORK STATE MUSEUM 


Sandy fields and thickets, Staten Island and Pennsylvania, south to 
Georgia. Flowering in late May and June. In the southern states several 
additional species of Leather Flower (Viorna) are found, but this is the 
only one which enters New York. 


Barberry Family 
Berberidaceae 
Blue Cohosh 


Caulophyllum thalictroides (Linnaeus) Michaux 


Plate 75 

Stems erect, glabrous and glaucous when young, 1 to 3 feet high from 
a thickened, perennial rootstock; the base of the stem with two or three 
large, sheathing bracts, near the top of the stem a single, large, triternate, 
nearly sessile leaf and usually a similar but smaller leaf near the base of 
the inflorescence. Leaflets thin, oval, oblong or obovate, 1 to 3 inches 
long when mature, usually only partly developed at flowering time, three 
to five-lobed at the apex. Flowers several in a loose terminal panicle, 
greenish purple, one-quarter to one-half of an inch bread; sepals six, oblong; 
petals six, smaller, cucullate and opposite the sepals; stamens six. Each 
flower contains a single pistil with two ovules, which ripens into a globose, 
blue, glaucous, berrylike fruit, about one-third of an inch in diameter. As 
the seed grows it ruptures the thin, transparent pericarp before maturity. 

In woods and thickets, New Brunswick to South Carolina, west to 
Manitoba, Tennessee, Nebraska and Missouri. Flowering in April 
and May. 

May Apple; Wild Mandrake 


Podophyllum peltatum Linnaeus 


Plate 76 
Stems erect, 1 to 14 feet high, from a perennial, horizontal, poisonous 
rootstock. Basal leaves centrally peltate, often nearly a foot in diameter, 
long petioled, deeply five to nine-lobed, glabrous or pubescent and light 
green on the lower surface, darker above; lobes two-cleft and toothed at 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 73 


arg 
re 


VIRGIN’S BOWER; WOODBINE; WILD CLEMATIS 
Clematis virginiana 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 74 


ERECT SILKY LEATHER FLOWER 
Viorna ochroleuca 


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WILD FLOWERS OF NEW YORK IIl5 


the apex. Flowering stems appearing from different rootstocks, bearing 
one to three, usually two, similar leaves (rarely leaflets). Flowers 14 to 2 
inches broad, white, fragrant, on stout, nodding peduncles one-half to 2 
inches long, appearing from the base of the upper leaf or usually from the 
fork between the two leaves; sepals six, petallike and soon falling. Petals 
six to nine, flat, obovate, longer than the sepals; stamens twice as many as 
the petals. Ovary ovoid, forming in fruit a large, yellowish, ovoid, edible 
berry, 15 to 2 inches long, the numerous seeds inclosed in fleshy arils within 
the fruit. 

In low woods, moist banks and clearings, western Quebec and southern 
Ontario to Minnesota, Kansas, Florida, Louisiana and Texas. Flowering 
in May and June. 

The fruit is edible and harmless, although somewhat insipid and to 
many people its taste is disagreeable. Both foliage and root are said to 
be poisonous and serious results have followed the use of the leaves as 
greens. The root is a viclent purgative, resembling jalap in its action. 
Its popular name, Mandrake, relates it in no way to the Mandrake or 
Mandragora of the ancients and, notwithstanding its poisonous character 
it is a very respectable herb in comparison with the traditions of the 
Mandrake of the ancients, described as flourishing best under a gallows, 
with a root resembling a man in shape, uttering terrible shrieks when it 
was torn from the ground, and possessing the power of transforming men 
and beasts. 

Twin-leaf 
Jeffersonia diphylla (Linnaeus) Persoon 
Figure XVIII 

A smooth, perennial, fibrous-rooted plant, 6 to 8 inches high when in 
flower, later becoming 10 to 18 inches high. Leaves and flowering stems 
arising from a scaly base. Leaves glaucous beneath, long petioled, cordate 
or reniform, 3 to 6 inches long, 2 to 4 inches wide when mature, parted 
longitudinally into two obliquely ovate, blunt, lobed or entire divisions; 
lobes rounded with sinuses sometimes three-fourths of an inch deep. At 


116 NEW YORK STATE MUSEUM 


(Photograph by G. A. Bailey) 


Figure XVIII 
Twin-leaf 


(Jeffersonia diphylla (Linnaeus) Persoon) 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 75 


BLUE COHOSH 
Caulophyllum thalictroides 


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WILD FLOWERS OF NEW YORK WIL 7/ 


flowering time the leaves are but partially developed. Flowering stems 
without leaves and one-flowered. Flowers white, about I inch broad; 
the calyx with four (sometimes three or five) caducous, petallike 
divisions. Petals eight, flat, oblong, longer than the sepals. Stamens 
eight with slender filaments. Fruit a short-stalked capsule about 1 
inch long, opening at maturity near the summit by a half-circumscissle 
cleft. 

In moist woods, New York, Ontario and Pennsylvania to Wisconsin, 
Iowa, Virginia and Tennessee. Flowering in April and May. 


Poppy Family 
Rapa vera ec eae 
Bloodroot; Puccoon-root 
Sanguinaria canadensis Linnaeus 
Plate 77 

Leaves and stems glabrous, especially when young, from a stout, hori- 
zontal rootstock, one-half to 1 inch thick, densely clothed with thick, fibrous 
roots. Juice of the roots and stems red. Leaves basal on petioles 6 to 14 
inches long, palmately five to nine-lobed, the lobes repand or cleft at the 
apex and palmately veined. Flowering scapes, one-flowered, at length 
overtopped by the mature leaves, but at flowering time longer than the 
partially unfolded immature leaves. Flowers white or sometimes pinkish, 
1 to 1 inches broad; sepals two, soon falling. Petals eight to sixteen, 
oblong-spatulate, arranged in two or three rows and soon falling; stamens 
numerous, yellow. Fruit a narrow, one-celled, two-valved capsule, pointed 
at both ends, about 1 inch long. 

In rich woods and on shaded banks, Nova Scotia to Manitoba and 


Nebraska, south to Florida, Alabama and Arkansas. Flowering in April 
and May. 


118 NEW YORK STATE MUSEUM 


Fumewort Family 
Fumariacede 
Dutchman’s-breeches 
Bicuculla cucullaria (Linnaeus) Millspaugh 
Plate 78 

A rather delicate, smooth and somewhat tufted herbaceous plant from 
a bulbous, perennial base. Leaves all basal, numerous and slender-stalked, 
pale beneath, finely divided into many narrow segments. Flowers nodding, 
fragrant, few or several on a slender stalk which rises above the leaves; 
each flower about one-half to two-thirds of an inch long, and somewhat 
broader than long across the spreading basal spurs, white or faintly pink, 
yellow at the summit; the four petals in two pairs, the outer pair oblong, 
concave, each with a divergent spur at the base and the tip spreading, the 
inner pair narrow and minutely crested. Fruit an oblong pod, opening 
into two parts to the base when mature. 

In rich woods, Nova Scotia to North Carolina, west to Minnesota, 
Kansas and Missouri. Flowering in April and May. 

The Squirrel Corn or Turkey Corn (Bicuculla canadensis 
(Goldie) Millspaugh) is similar, but the spurs of the two outer petals are 
shorter, rounded and not divergent, the inner pair of petals is conspicu- 
ously crested and the roots have numerous small tubers. The Wild 
Bleeding Heart (Bicuculla eximia (Ker) Millspaugh) of the 
western part of the State has pink flowers. 


Mountain Fringe; Alleghany Vine 
Adiumia fungosa (Aiton) Greene 
Plate 79b 
Stems weak, slender, climbing several feet over other plants by its 
slender petioles from a biennial root. Leaves two to three-pinnate, the 
leaflets slender stalked, lobed or entire, very thin, ovate or cuneate, about 


one-fourth of an inch long and pale beneath. Flowers numerous in axillary 


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WILD FLOWERS OF NEW YORK II9 


drooping cymes; petals four, united into a narrowly ovate-cordate, spongy 
corolla about one-half of an inch long and one-fourth of an inch broad 
at the base, four-lobed at the apex, greenish purple, or pink, usually rather 
persistent, dry and brownish with age; stamens six, united below and 
adherent to the petals. Fruiting capsule oblong, included in the persistent, 
dry corolla. 

In moist woods, thickets and shaded cliffs, New Brunswick to Ontario 
and Michigan, south to North Carolina and Tennessee. Flowering from 
June to October. Frequent in cultivation. 


Pink or Pale Corydalis 
Capnoides sempervirens (Linnaeus) Borkhausen 
Plate 80 

Stems and leaves glabrous, glaucous, erect or ascending, 6 inches to 
2 feet high and freely branching. Lower leaves I to 5 inches long, short 
petioled, the upper leaves sessile or nearly so, all divided into numerous 
obovate or cuneate segments, toothed or entire, obtuse. Flowers numerous, 
panicled, borne in cymose clusters at the ends of the branches. Each 
flower one-half to two-thirds of an inch long, pink or rarely white with a 
yellow tip; sepals two, small; corolla irregular, deciduous; petals four, 
erect-connivent, one of the outer pair with a spur at the base about one- 
eighth of an inch long, the inner pair narrower, keeled at the back. Cap- 
sules narrowly linear, erect, I to 2 inches long. 

In rocky piaces, Nova Scotia to Alaska, Georgia, Minnesota, Montana 
and British Columbia. Flowering from May to September. 

Two other species of this genus are occasionally found in this State, 
both with low, diffusely spreading stems and with yellow flowers. They are 
the Yellow Corydalis (Capnoides flavulum (Rafinesque) Kuntze) 
with flowers about one-fourth of an inch long and short spurred; and the 
Golden Corydalis (C. aureum (Willdenow) Kuntze) with flowers 
slightly more than one-half of an inch long and spurs one-half the length 
of the corolla. 


120 NEW YORK STATE MUSEUM 


Mustard Family 
Cruciate xrare 
Lyre-leaved Rock Cress 
Arabis lyrata Linnaeus 
Plate s4b 
A low, tufted perennial or biennial plant with ascending or erect stems, 
4 to 12 inches high, smooth above, pubescent below or glabrous throughout. 
Basal leaves lyrate-pinnatifid, 1 to 2 inches long, spatulate or oblanceolate, 
pubescent or glabrous; stem leaves entire or toothed, spatulate or linear, 
one-half to 1 inch long. Flowers white, one-fourth of an inch broad 
or less, several or numerous in a terminal raceme which elongates in 
fruit; pedicels ascending, about one-third of an inch long or longer in fruit; 
petals four, much longer than the four stamens. Fruit a linear, slightly 
flattened pod three-fourths to 1} inches long; seeds in one row, oblong 
and wingless. 
Rocky and sandy places, Connecticut and Ontario west to Manitoba 
and Alaska, south to Virginia, Tennessee, Missouri and British Columbia. 


Flowering from April to September. 


Bulbous Cress; Cuckoo-flower 


Cardamine bulbosa (Schreber) Britton, Sterns & Poggenberg 
Plate goa 

A slender, erect, herbaceous plant with smooth stems, 6 to 20 inches 
high from a perennial, tuber-bearing root, simple or rarely branched. 
Leaves of two sorts, those of the stem sessile and clasping or the lower ones 
very short petioled, rather distant from one another, oblong or lanceolate, 
blunt, toothed or entire, 1 to 2 inches long; the basal leaves oval or 
orbicular, one-half to 2 inches broad, often slightly heart-shaped, toothed 
or usually entire, with long, slender petioles. Flowers white, about one- 
half of an inch broad, in a terminal, several to many-flowered cluster; 


petals four, three to four times the length of the four sepals. Fruit a 


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PINK OR PALE CORYDALIS 
Capnoides sempervirens 


WILD FLOWERS OF NEW YORK 121 


number of erect, very slender pods, narrowed at each end and about an 
inch long. 

In low, wet woods, thickets and wet meadows, New Brunswick to 
Vermont, southern Ontario and Minnesota, south to Florida and Texas. 
Flowering in April and May, or in cool woods as late as early June. 


The Purple Cress (Cardamine douglassii (Torrey) Britton) 
has leaves which are more angularly toothed and showy purple flowers 
often three-fourths of an inch broad. It is found in cold, springy places 
in the northern part of the State. 

The: “rue Water-cress (Sisymbrium nastuntium-aqiua- 
ticum Linnaeus) is a native of Europe, but is common in brooks and 
streams in most parts of New York. 


Cut-leaved Toothwort or Pepperroot 
Dentaria laciniata Muhlenberg 
Plate 81a 

Stems erect, pubescent or glabrous, 8 to 15 inches high from a deep, 
perennial, tubercled, jointed rootstock, the joints easily separable. Leaves 
all petioled, 2 to 5 inches broad, those on the stem usually three and forming 
a whorl, rarely distant, three-parted nearly to the base, the divisions lanceo- 
late, linear or oblong, the lateral ones often deeply cleft, all deeply toothed 
or lobed; basal leaves similar, usually developing later than the flowering 
stems. Flowers numerous in a stout, broad raceme, two-thirds to three- 
fourths of an inch broad, pink or white. The four petals longer than 
the sepals. Stamens six. Fruiting pod linear, ascending, 1 to 13 inches 
long. 

In moist or rich woods, Quebec to Florida, west to Minnesota, Kansas 
and Louisiana. Flowering in April and May. 


Two-leaved Toothwort or Crinkleroot 
Dentaria diphylla Michaux 
Plate 81b 
Stems stout, simple, glabrous, 6 to 14 inches high from a perennial, 
notched but continuous rootstock. Basal leaves long petioled, 4 to 5 inches 


[22 NEW YORK STATE MUSEUM 


broad, with three broadly ovate, dentate or somewhat lobed leaflets each 
about 2 inches long; stem leaves usually two, opposite or nearly so, short 
petioled and also three-divided, the leaflets often narrower than those 
of the basal leaves. Flowers white, one-half to two-thirds of an inch 
broad. 

In rich woods, Nova Scotia and New Brunswick to Minnesota, south 
to South Carolina and Kentucky. Flowering usually in late April and 
in May. 

The Large Toothwort (Dentaria maxima Nuttall) has a 
jointed rootstock and three stem leaves (sometimes two or as many as five 
or six), alternate, with ovate and obovate toothed and cleft leaflets and 
large pale-purple flowers. It is rare and local in distribution. 

The species of Dentaria are members of the Mustard family (Cruciferae) 
which contains a very large number of small-flowered, inconspicuous plants, 
many of them weeds, as well as a number of cultivated species which have 
become naturalized or established throughout the State. 


American Sea Rocket 
Cakile edentula (Bigelow) Hooker 
Plate 82a 

Plant very fleshy throughout, bushy branched from a deep, annual root, 
the lower branches spreading or ascending, the center ones erect, a few 
inches to a foot high. Leaves oblanceolate or obovate, obtuse, lobed or 
toothed, narrowed at the base, the lower leaves 2 to 5 inches long, the 
upper leaves smaller. Flowers light purple, less than one-fourth of an inch 
broad, the four petals long-clawed, more than twice the length of the sepals; 
fruit one-half to 1 inch long, the upper joint slightly longer than the lower, 
ovoid, angled, flattened, narrowed above into a beak; lower joint obovoid, 
not flattened. 

Sandy places along the seashore, Newfoundland to New Jersey and 


Florida, and along the Great Lakes, New York to Minnesota. 


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A. AMERICAN SEA ROCKET 
Cakile edentula 


B. YELLOW MOUNTAIN SAXIFRAGE 
Leptasia aizoides 


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WILD FLOWERS OF NEW YORK I 


Pitcher Plant Family 
Sid) er arCremala Gierare 
Pitcher Plant; Sidesaddle Flower 
Sarracenia purpurea Linnaeus 
Plate 83 

Leaves tufted, ascending, hollow, much inflated and trumpet-shaped, 
4 to 12 inches long, with a broad, lateral wing and an erect terminal lid or 
lamina, glabrous except the inner side of the lamina and the inner surface 
of the pitchers, which are densely clothed with stiff, reflexed hairs, purple- 
veined or sometimes green, yellowish or reddish all over, narrowed into 
petioles below, persistent over winter. Roots large, stout and fibrous. 
Flowers solitary on slender scapes, I to 2 feet high, nodding, deep purple 
or rarely yellow, nearly globose, 14 to 2} inches broad; sepals five, green, 
with three or four bracts at the base; petals five, obovate, narrowed in the 
middle, incurved over the yellowish style. Style dilated at the apex into 
a peltate umbrellalike structure with five rays which terminate under its 
angles in hooked stigmas. 

In peat bogs and wet sphagnous places, Labrador to the Canadian 
Rocky mountains, Florida, Kentucky and Iowa. Flowering in May and 
June. The pitcher-shaped leaves usually contain more or less water in 
which are numerous drowned insects which furnish food for the larvae of 
a fly which is instrumental in the cross-pollination of the flowers. 


Sundew Family 
Dts OISie Tai ce ane 
Spatulate-leaved Sundew 
Drosera intermedia (Linnaeus) Hayne 
Plate 84 

A tufted bog plant with erect, flowering scapes, 2 to 8 inches high, 
and elongated rootstocks. Leaf blades ascending, spatulate, obtuse at the 
apex, one-fourth to three-fourths of an inch long, one-half to one-third as 


124 NEW YORK STATE MUSEUM 


wide as long, clothed above with long, glandular hairs secreting a fluid 
which entraps insects, narrowed below into glabrous petioles one-half to 
13 inches long; usually the entire foliage reddish or greenish red in color. 
Flowers several in one-sided racemes; petals five, white, slghtly longer than 
the greenish sepals; the one-celled ovary surmounted by three styles, each 
deeply two-parted so as to appear like six. 

In bogs and sphagnous places, Newfoundland to Saskatchewan, south 
to Florida and Louisiana, and also in northern Europe. Flowering from 
June to August. 

The Spatulate-leaved Sundew is not so common as the Round-leaved 
Sundew (Drosera rotundifolia Linnaeus), with orbicular leaf 
blades. Two additional Sundews occur in New York, namely the Oblong- 
leaved Sundew (Drosera longifolia Linnaeus), with leaf blades 
elongated-spatulate, six to eight times as long as wide; and the Thread- 
leaved Sundew (D. filiformis (Linnaeus) Rafinesque), with linear 
leaves ten to fifteen times as long as wide and purple flowers. The last 
grows in wet sand near the coast, the others in bogs. 


Virginia Stonecrop Family 
Penthoraceaec 
Ditch or Virginia Stonecrop 
Penthorum sedoides Linnaeus 
Plate 87b 
Stems erect, glabrous, often branched and angled above, 6 inches 
to 2 feet high, from a perennial root. Leaves alternate, sessile, lanceolate 
or narrowly elliptic, acuminate at each end, finely toothed, 2 to 4 inches 
long, one-half to 1 inch wide. Flowers perfect, yellowish green, in two 
or three forked, one-sided cymes, the branches 1 to 3 inches long. Each 
flower about one-fifth of an inch broad; calyx five-parted, the sepals 
triangular-ovate, pointed, shorter than the flattish capsule; stamens ten; 
petals often lacking, when present, linear or linear-spatulate. Fruit a 
depressed, five-lobed capsule with five divergent tips. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 


Plate 83 


PITCHER PLANT; SIDESADDLE FLOWER 
Sarracenia purpurea 


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WILD FLOWERS OF NEW YORK 125 


Common in ditches and swampy places, New Brunswick to Florida, 
west to Nebraska, Kansas and Texas. Flowering from July to September. 
Not a very attractive plant, but the only American representative 
of the Penthoraceae (Virginia Stonecrop family), which is joined with 
the Saxifrage family by some authors and to the Orpine family by others. 


Grass-of-Parnassus Family 
IP iri 2,9 Si CCA 
Carolina Grass-of-Parnassus 


Parnassia caroliniana Michaux 
Plate 85 


Flowering scape 6 to 20 inches high, with a sessile, ovate, clasping 
leaf below the middle. Basal leaves very numerous, ovate, oval, or nearly 
orbicular, obtuse at the apex, rounded or slightly cordate at the base, or 
decurrent on the petiole, 1 to 2 inches long, on petioles 2 to 6 inches long. 
Flowers three-fourths to 1} inches broad; calyx lobes five, ovate-oblong, 
obtuse and much shorter than the five broadly oval, white, greenish veined 
petals; each petal with a set (usually three) of gland-tipped staminodia at 
the base which do not exceed the five fertile stamens in length, the latter 
alternate with the petals. Fruit a one-celled capsule about one-half of an 
inch long. 

In swamps, low meadows and boggy places, New Brunswick to Mani- 
toba, south to Virginia, Illinois and Iowa. Flowering from July to Sep- 
tember, rarely earlier than July in our latitude. At Taberg growing on wet 
cliffs with the Yellow Mountain Saxifrage and Dwarf Canadian Primrose. 


Saxifrage Family 
paxifragaceae 
Yellow Mountain Saxifrage 
Leptasea aizoides (Linnaeus) Haworth 
Plate 82b 
Stems tufted, forming loose or dense leafy mats, 2 to 7 inches high. 
Leaves alternate, linear, thick, fleshy, sharply pointed at the apex, sessile, 


126 NEW YORK STATE MUSEUM 


one-third to three-fourths of an inch long, one-eighth of an inch wide or 
usually less, sometimes sparingly ciliate on the margins. Flowers several, 
corymbose, one-third to two-thirds of an inch broad on slender pedicels; 
petals five, oblong, yellow and often spotted with orange, longer than 
the ovate calyx lobes and alternate with them. Stamens ten; ovary almost 
superior, the two capsules united to above the middle. 

On wet or dripping rocks, Newfoundland and Labrador to Vermont, 
northern and western New York, and west through Arctic America to the 
Rocky mountains. Also in Alpine and Arctic Europe and Asia. Flowering 
in July and August. 

Early Saxifrage 
Micranthes virginiensis (Linnaeus) Small 
Plate 87a 

Flowering stem 4 to 12 inches high, viscid-pubescent with whitish 
hairs, leafless or with a few green bracts at the base of the inflorescence. 
Leaves all basal, 1 to 3 inches long, obovate, or oval, toothed, blunt or 
pointed at the apex, spatulate at the base and narrowed into a margined 
petiole; inflorescence cymose, becoming paniculate by the elongation of 
the lower branches. Flowers white, less than one-fourth of an inch broad; 
calyx of five erect, triangular, pointed lobes; petals five, oblong-spatulate, 
obtuse, longer than the calyx; stamens ten, carpels of the fruit usually 
two, nearly separate, widely divergent when mature. 

In dry or rocky woodlands, banks and ledges, New Brunswick to 
Minnesota, south to Georgia and Tennessee. Flowering from April until 
June. 

Swamp Saxifrage 
Micranthes pennsylvanica (Linnaeus) Haworth 
Plate 86 

Flowering scape stout, viscid-pubescent, 1 to 35 feet high, with green 
bracts at the inflorescence, otherwise leafless. Leaves all basal, large, 
oval, ovate, obovate or oblanceolate, pubescent or nearly glabrous, 4 to 
10 inches long, obtuse at the apex, narrowed below into a broad petiole, 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 85 


GRASS-OF-PARNASSUS 
Parnassia caroliniana 


= 
— 


WILD FLOWERS OF NEW YORK 127 


the margins denticulate or repand. Flowers in elongated, loose, terminal 
panicles, greenish. Each flower about one-eighth of an inch broad or 
slightly broader; the obtuse calyx lobes reflected; petals five, lanceolate 
or linear-lanceolate, twice as long as the calyx. 

Swamps, wet banks and wet woods, Maine to Ontario, Minnesota, 
Virginia, Iowa and Missouri. Flowering in May and June. 


Foamflower; False Miterwort 
Tiarella cordifolia Linnaeus 
Plate 88 
Flowering scapes 6 to 12 inches high, slender and pubescent, from 
a rather stout, perennial root. Leaves all basal, long petioled, broadly 
ovate or nearly orbicular, cordate at the base, three to seven-lobed, blunt 
or pointed at the apex, 2 to 4 inches long, margins crenate or dentate, 
pubescent above with scattered hairs, glabrate or downy along the veins 
beneath. Flowers white, forming a terminal raceme, I to 4 inches long. 
Each flower about one-fourth of an inch broad; petals five, oblong, entire 
or slightly toothed, somewhat longer than the five white calyx lobes; stamens 
ten, anthers reddish or yellowish. Fruit of two very unequal carpels, about 
one-fourth of an inch long, reflexed on slender pedicels. 
In rich, moist woods, Nova Scotia to Ontario and Minnesota, south 
along the mountains to Georgia, and west to Indiana and Michigan. 
Flowering from April to June. Also known as Coolwort. 


Alumroot 
Heuchera americana Linnaeus 
Plate 89 
Flowering stem rather stout, 13 to 3 feet high, leafless, glandular, 
hirsute. Leaves basal, long petioled, 3 to 4 inches wide with seven to 
nine rounded, crenate-dentate lobes; the older leaves glabrous or with 
scattered hairs on the upper surfaces, new leaves usually somewhat pubes- 
cent. Flowers greenish yellow, in elongated panicles; calyx tube broadly 


128 NEW YORK STATE MUSEUM 


campanulate, nearly regular, somewhat less than one-fourth of an inch 
long, five-lobed; petals five, very small, greenish and alternate with the 
lobes of the calyx which they do not exceed in length. Stamens five, 
projecting out from the calyx more than one-half their length, anthers 
orange. 

In dry or rocky woods and banks, Ontario to Connecticut, west to 
Minnesota, south to Alabama and Louisiana. Flowering from May to 
August. 

Two-leaved Bishop’s Cap; Miterwort 
Mitella diphylla Linnaeus 
Plate 90b 

Stems erect, often several together from a perennial root, 8 to 17 
inches high, pubescent, each stem bearing a pair of opposite, sessile or 
nearly sessile leaves near or above its middle. Basal leaves broadly ovate, 
cordate at the base, acute or long pointed at the apex, three to five-lobed, 
toothed, rather rough-hairy on both sides, 1 to 2 inches long. Flowers 
small, white, rather distant from one another, in a very narrow, elongated, 
erect raceme, 3 to 8 inches long; calyx tube bell-shaped, five-lobed; petals 
five, finely pinnatifid. Fruiting capsules one-celled, two-valved at the 
apex, many seeded, somewhat flattened and broad, seeds smooth, black 
and shiny. 

In rich woods, Quebec to Minnesota, North Carolina and Missouri. 
Flowering in April and May. 

In cold woods and bogs of the northern part of the State occurs a 
smaller species of Miterwort, Mitella nuda Linnaeus, with reniform- 
orbicular, basal leaves and the stems usually without leaves; flowers 
greenish yellow. Another species, M. oppositifolia Rydberg, has 
been described from central New York which differs from M. diphylla 
only in having long-petioled stem leaves, lanceolate calyx lobes and filiform 


divisions to the petals. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum = Plate 86 


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Tiarella cordifolia 


WILD FLOWERS OF NEW YORK 129 


Rose Family 
Rosaceae 
Meadowsweet; Quaker Lady 
Spiraea latifolia (Aiton) Borkhausen 
Plate 93a 

An erect shrub, 2 to 6 feet high, usually more or less branched above 
and smooth with reddish stems. Leaves short petioled, blades oblanceolate 
or obovate, glabrous or nearly so, coarsely toothed, 1 to 2 inches long, one- 
half to 13 inches wide, usually larger on young shoots, obtuse or slightly 
pointed at the apex, rounded or tapering at the base, pale beneath. Flowers 
white or pinkish, in dense terminal panicles, each flower about one-fourth 
of an inch broad or less; petals four or five, inserted on the calyx; stamens 
numerous. Pistils commonly five, alternate with the calyx lobes. 

In moist or rocky places, in open woods, or in old meadows and along 
roadsides, Newfoundland to Saskatchewan, Virginia and western Penn- 
sylvania. Flowering from June to August. 

The Narrow-leaved Meadowsweet (Spiraea alba Du Roi) has 
yellowish brown branches, narrowly oblanceolate to oblong leaves and 
white flowers. It is much less abundant than Spiraea latifolia, 
and is found in wet soil, Ontario to New York, south to North Carolina, 


west to Indiana, Missouri and Saskatchewan. 


Hardhack; Steeplebush 


Spiraea tomentosa Linnaeus 


Plate 94a 
Erect, shrubby and perennial at least below, the tops usually dying 
back, 1 to 3 feet tall, usually simple; stems floccose-pubescent. Leaves 
short petioled, ovate or oval, 1 to 2 inches long, one-half to 1 inch wide, 
unequally toothed, blunt or pointed at the apex, narrowed or rounded 
at the base, smooth and dark green above, woolly-pubescent with whitish 
hairs beneath. Flowers pink or purplish, rarely white, in dense terminal 


130 NEW YORK STATE MUSEUM 


panicles, each flower about one-fifth of an inch broad; divisions or follicles 
of the fruit pubescent. 

In wet meadows, swamps and low ground, Nova Scotia to Manitoba 
south to Georgia and Kansas. Flowering from July to September. 

The Corymbed Spiraea (Spiraea corymbosa Rafinesque), 
with oval, ovate or orbicular leaves, slightly cordate at the base, or rounded, 
and with small white flowers in dense terminal, often leafy corymbs, is 
found occasionally on rocky banks and in wood of the southern part of 
the State. 

Indian Physic; Bowman’s Root 
Porteranthus trifoliatus (Linnaeus) Britton 
Plate 91 

Stems erect, herbaceous, 2 to 4 feet high, from a perennial root, usually 
branched, glabrous or slightly pubescent. Stipules small, one-eighth 
to one-fourth of an inch long, entire or toothed. Leaves sessile or nearly 
so, three-foliate; leaflets oval, ovate, lanceolate or slightly obovate, long 
pointed at the apex, narrowed at the base, 2 to 3 inches long, irregularly 
toothed. Flowers white or pinkish, one-half to two-thirds of an inch 
long on slender peduncles, in loose, terminal, leafy panicles; calyx reddish, 
five-toothed, ten nerved; petals five, linear-lanceolate, somewhat unequal. 

In woods and thickets, Ontario and New York to Michigan, Georgia 
and Missouri. Flowers in June and July. 

A. closely related species; Porteranthus -stipulaitus 
(Muhlenberg) Britton, has incised leaflets, broad, foliaceous, incised 
stipules and slightly smaller flowers. Its range is about the same. 


Common Five-finger or Cinquefoil 
Potentilla canadensis Linnaeus 
Plate 92a 
Stems ascending, a few inches high, spreading by slender runners 
3 inches to I or 2 feet long; the pubescence of the stem, petioles and peduncles 
spreading. Leaves petioled, digitately five-foliated; leaflets oblanceolate 


Memoir 15 


WILD FLOWERS OF NEW YORK 


N. Y. State Museum 


Plate 89 


ALUMROOT 
Heuchera americana 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 90 


A. BULBOUS CRESS; CUCKOO-FLOWER B. TWO-LEAVED BISHOP’S CAP OR MITERWORT 
Cardamine bulbosa Mitella diphylla 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 91 


INDIAN PHYSIC; BOWMAN’S ROOT 
Porteranthus trifoliatus 


WILD FLOWERS OF NEW YORK 131 


to oblong, blunt at the apex, narrowed at the base, one-half to 1 inch long, 
toothed. Flowers yellow, one-fourth to three-fourths of an inch broad 
on slender, axillary, one-flowered peduncles; the first flower appearing 
from the axil of the second stem leaf; petals five, broadly oval, slightly 
longer than the acute calyx lobes and the linear-lanceolate bractlets; 
stamens about twenty. 

In dry soil of meadows and fields, New Brunswick to Georgia, 
Minnesota and Texas. Flowering from April to August. 

The genus Potentilla contains a number of representatives in New 
York, several of which are introduced species. Of these, the most common 
is the Rough Cinquefoil (Potentilla monspeliensis Linnaeus), 
with stout, erect, hairy stems and leaves, and yellow flowers in which the 
calyx lobes exceed the petals in length. 

The Silvery Cinquefoil (Potentilla argentea Linnaeus) is a 
native species of dry soils or rocky places, with spreading or ascending, 
white, woolly-pubescent stems, leaflets green and smooth above and white- 
tomentose beneath, the margins revolute, and small yellow flowers. 


Rough-fruited Cinquefoil 
Potentilla recta Linnaeus 
Plate 95 
Stems erect, stout, branched above, villous-pubescent, I to 2 feet 
high from a stout, perennial root, with ovate-lanceolate stipules, the 
lower foliaceous and cut. Leaves digitately five to seven-foliate, petioled 
or the upper leaves nearly sessile; leaflets oblanceolate, blunt at the apex, 
narrowed at the base, pubescent with scattered hairs above, more densely 
pubescent beneath, sharply toothed, 1 to 3 inches long. Flowers numerous, 
bright or dull yellow, about three-fourths of an inch broad, in terminal, 
cymose clusters; stamens about twenty. 
Roadsides, fields and waste places, Maine to Ontario, New York, 
Virginia and Michigan. Naturalized from Europe and Asia. Flowering 
from June to September. 


132 NEW YORK STATE MUSEUM 


Silverweed; Wild or Goose Tansy 
Argentina anserina (Linnaeus) Rydberg 
Plate 92b 

Stems and leaves herbaceous and tufted from a perennial root; spreading 
by slender runners, 8 inches to 3 feet long. Leaves pinnate, 3 to 18 inches 
long; leaflets seven to twenty-five, oblong, oblanceolate or obovate, obtuse, 
the lower leaflets usually smaller, often with still smaller ones interspersed, 
all sharply toothed, nearly glabrous above, white or silky-pubescent beneath. 
Flowers yellow, three-fourths to 1§ inches broad, solitary on erect axillary 
peduncles; petals five, broadly oval or obovate, longer than the five ovate, 
pointed sepals and the five oval bractlets; stamens about twenty, borne 
around the base of the hemispheric, villous receptacle. 

Lake shores, sandy fields and salt meadows, New Jersey to Greenland, 
west to Nebraska, British Columbia and Alaska, south in the Rocky 
mountains to New Mexico and California; also in Europe and Asia. 
Flowering from May to September. Consists of several or numerous 
races, differing in the size and shape of the leaflets, and slightly in the 
achenes. Small northern plants have been referred to A.egedii of 
Greenland, and a form from Oneida lake has been described as A. ba b- 


cockiana Rydberg. 


Purple or Marsh Cinquefoil; Purplewort 
Comarum palustre Linnaeus 
Plate 96a 
Stems decumbent, often rather long, somewhat woody and perennial 
at the base; pubescent above. Leaves pinnate, upper leaves three to 
five-foliate, nearly sessile, lower ones successively longer petioled, five 
to seven-foliate; leaflets oblong or oval, sharply toothed, blunt or pointed 
at the apex, narrowed at the base, 1 to 3 inches long; petioles sheathed 
at the base by large membranous stipules. Flowers conspicuous, three- 
fourths to 14 inches broad; calyx deeply five-lobed with five narrow bractlets, 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 92 


A. COMMON FIVE-FINGER OR CINQUEFOIL 
Potentilla canadensis 


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A. HARDHACK; STEEPLEBUSH B. AMERICAN GREAT BURNET 
Spiraea tomentosa Sanguisorba canadensis 


WILD FLOWERS OF NEW YORK 133 


the calyx lobes ovate, acuminate, red or purple within, much exceeding 
the purple, ovate-lanceolate petals; pistils numerous, seated on an enlarged, 
pubescent receptacle which becomes spongy in fruit. 

In swamps and peat bogs, Greenland and Labrador to New Jersey, 
Iowa, British Columbia, Wyoming, Alaska and California; also in northern 
Europe and Asia. Flowering from June to August. 


Wild or Scarlet Strawberry 
Fragaria virginiana (Linnaeus) Duchesne 
Plate 97a 

Plants tufted, usually several or many together, dark green, foliage 
villous-pubescent with spreading hairs. Leaves thick, with three broadly 
oval or obovate, coarsely toothed leaflets, the terminal one usually narrowed 
at the base; petioles 2 to 6 inches long. Flowering scapes as long or shorter 
than the leaves, bearing several white flowers on appressed-pubescent 
pedicels. Flowers one-half to three-fourths of an inch broad, white, petals 
obovate. Fruit red, ovoid, the achenes imbedded in pits. 

In rather dry soil, fields, hillsides etc., Newfoundland to South Dakota, 
Florida and Oklahoma. Flowering in April and May. 

The European Wood Strawberry (Fragaria vesca Linnaeus) is 
a common escape everywhere in the east and frequently hybridizes with 
F. virginiana, so that some forms are difficult to classify. The 
American Wood Strawberry (Fragaria americana _ (Porter) 
Britton), with longer flowering scapes and elongated-conic fruit, with 
achenes borne on its shining, even surface and but slightly attached to it, 
is a common species in rocky woodlands. The Northern Wild Strawberry 
(Fragaria canadensis Michaux) has a long, slender fruit with 
the achenes sunk in pits and oblong or narrowly obovate leaflets. It ranges 
across the northern part of the state. 


134 NEW YORK STATE MUSEUM 


Shrubby Five-finger or Cinquefoil 
Dasiphora fruticosa (Linnaeus) Rydberg 
Plate 96b 

A branching shrub with ascending or erect, leafy branches, 1 to 4 feet 
high. Leaves pinnate, leaflets five to seven, linear-oblong or oblanceolate, 
pointed at each end, one-half to 1 inch long, silky-pubescent, the margins 
entire and revolute; stipules membranous, ovate-lanceolate, entire and 
pointed. Flowers terminal, in dense or loose cymose clusters, or solitary, 
bright yellow, three-fourths to 1} inches broad; the five calyx lobes ovate, 
with five bractlets; petals five, nearly orbicular and longer than the calyx 
lobes. Achenes of the fruit covered with long straight hairs. 

In swampy or marshy places, often in moist rocky places, Labrador and 
Greenland to Alaska, south to New Jersey, Illinois, Minnesota, Rocky 
mountains and California; also in northern Europe and Asia. In northern 
New England it is often a troublesome, bushy weed. Flowering from 
June to September. 

American Great Burnet 
Sanguisorba canadensis Linnaeus 
Plate o4b 

An erect, leafy, herbaceous plant from a thick, perennial root; stems 
sometimes decumbent at the base, glabrous or somewhat pubescent below, 
simple or branched above, I to 6 feet high. Leaves odd-pinnate, the lower 
leaves long petioled, sometimes 1 to 13 feet long; leaflets seven to fifteen, 
ovate, oblong or oval, blunt or pointed at the apex; pointed, blunt or even 
cordate at the base, sharply toothed. Flowers white, in dense terminal 
spikes, I to 6 inches long; petals none; calyx tube turbinate, constricted at 
the throat, four-winged, four-lobed, the lobes petallike, concave and 
deciduous; stamens four, their filaments long and white. 

In swamps and low meadows, sometimes in bogs, Newfoundland to 
Michigan, south to Georgia. Flowering from July to September or even 
later. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 95 


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WILD FLOWERS OF NEW YORK 135 


Common Agrimony 
Agrimonia striata Michaux 


(Agrimonia brittoniana Bicknell) 


Plate 98b 

Stem rather stout and usually with some straight, nearly erect branches, 
2 to 6 feet tall from a perennial fibrous root, pubescent with short, spread- 
ing, brownish hairs, somewhat appressed above. Leaves numerous, alternate, 
compound with seven to nine, or rarely eleven, oblique leaflets, tetragonal- 
elliptic to rhomboid-lanceolate, pointed and sharply toothed, rather thick 
and somewhat rough, dull green above, softly pubescent beneath, usually 
several pairs of small, interposed leaflets; stipules lanceolate, pointed and 
cut-toothed. Flowers numerous in long, erect or ascending racemes; each 
flower about one-fourth of an inch wide; petals five, bright yellow; calyx 
tube in fruit long-turbinate, about one-fourth of an inch long, deeply grooved, 
unmargined; the bristles numerous, often purplish, short, crowded, inflexed 
and connivent over the sepals. 

Thickets, open woods and roadsides, Newfoundland to Saskatchewan, 
south to West Virginia, Nebraska and Mexico. Flowering from June 


to September. 
Barren or Dry Strawberry 


Waldsteinia fragarioides (Michaux) Trattinnick 
Plate 09 

A perennial, herbaceous plant resembling a strawberry, with creeping, 
rather stout rootstock. Leaves tufted, mainly basal, long petioled, glabrous 
or somewhat pubescent, three-foliate; leaflets obovate, obtuse at the apex, 
tapering at the base with crenate or sometimes incised margins, I to 2 
inches long. Flowering scapes slender, erect, bracted, corymbosely three 
to eight-flowered; pedicels slender, often drooping; flowers yellow, one-half 
to two-thirds of an inch broad; petals five, obovate and longer than the 
five ovate-lanceolate, acute calyx lobes; stamens eight, inserted on the 
throat of the calyx; achenes of the fruit four to six, finely pubescent. 

Rocky woods, shaded hillsides and banks, New Brunswick to Ontario, 
Minnesota, Michigan, Indiana and Oregon. Flowering in May and June. 


126 NEW YORK STATE MUSEUM 


Yellow Avens; Camproot 
Geum strictum Aiton 
Plate 98a 
Stems erect or nearly so, pubescent, somewhat branched, 2 to 4 feet 
high, basal leaves lyrate-pinnate with five to seven obovate, cuneate, 
toothed or lobed leaflets, with a few smaller leaflets interspersed, terminal 
leaflets largest, broadly ovate or cuneate; stem leaves short petioled or 
sessile with three to five ovate or oblong, acute segments. Flowers 
yellow, several, terminal and short peduncled, one-half to three-fourths 
of an inch broad; petals obovate, exceeding the five calyx lobes. Stamens 
numerous. 
In low, shaded ground, swamps and wet meadows, Newfoundland 
to British Columbia, south to New Jersey, Pennsylvania, Missouri and 
New Mexico. Flowering from June to August. 


Purple or Water Avens 
Geum rivale Linnaeus 
Plate 100 
Stems simple or nearly so, erect, pubescent, 1 to 3 feet high, from 
stout perennial roots. Basal leaves lyrately and interruptedly pinnate, 
petioled, the lateral segments generally few and small, the terminal one 
to three leaflets much larger, all sharply and irregularly lobed and toothed; 
stem leaves rather far apart, short petioled or sessile, simple or three- 
foliated. Flowers few, terminal, purple or purplish, nodding, three-fourths 
to 1 inch broad; petals obovate emarginate, abruptly narrowed into a 
claw; calyx lobes spreading, purple; head of the fruit stalked in the calyx; 
achenes pubescent, style jointed, plumose below, about one-fourth of an 
inch long. 
In swamps and low grounds, Newfoundland to British Columbia, 
New Jersey, Pennsylvania, Michigan and Colorado; also in northern Europe 
and Asia. Flowering from early June until the latter part of July. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 98 
5 = 1 


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A, 


A. YELLOW AVENS; CAMPROOT B. COMMON AGRIMONY 
Geum strictum Agrimonia striata 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 99 


BARREN OR DRY STRAWBERRY 
Waldsteinia fragarioides 


WILD FLOWERS OF NEW YORK 137 


There are several other Avens in New York, all except G. rivale 
with reflexed calyx lobes. G. virginianum Linnaeus and G. cana- 
dense Jacquin have white flowers. G. vernum (Rafinesque) 
Torrey & Gray; G. strictum Aiton (described above), and G. 
meyerianum Rydberg, have yellow flowers. G.macrophyllum 
Willdenow, a boreal species found only in the Adirondacks, in this State, 
also has yellow flowers. For complete descriptions of these additional 
species the student should refer to Gray’s Manual or Britton and Brown’s 
Illustrated Flora. 


Purple-flowering Raspberry; Thimbleberry 
Rubus odoratus Linnaeus 
Plate ror 

Shrubby, erect and branched, perennial; new growth glandular-pubes- 
cent and somewhat bristly but not prickly, 3 to 5 feet high. Leaves simple, 
petioled, large, 5 to 10 inches broad, three to five-lobed, cordate at the base, 
pubescent, especially on the veins beneath, the lobes long pointed, the 
middle lobe usually longer than the others; flowers rather numerous in 
corymbose, terminal clusters, purple, 1 to 2 inches broad; calyx lobes tipped 
with long, slender appendages; petals five; fruit red when ripe, broad and 
thin, scarcely edible. 

In rocky woods and thickets, Nova Scotia to Ontario and Michigan, 
south to Georgia and Tennessee. Flowering from June to August. 

New York contains a large number of native raspberries and black- 
berries. The principal species of Raspberries are R. strigosus Michaux 
(red); R.neglectus Peck (purple), and R. occidentalis Linnaeus 
(Black Raspberry). The Blackberries are R.triflorus Richards (Dwarf 
Red Blackberry), R.canadensis Linnaeus (Northern Blackberry), 
R. allegheniensis Porter (Mountain Blackberry), R. argutus 
Link (Tall Blackberry), and also the Dewberries. See Britton and Brown’s 
Illustrated Flora for complete descriptions. 


138 NEW YORK STATE MUSEUM 


Dewberry; Low Running Blackberry 


Rubus procumbens Muhlenberg 
Plate 93b 


Stems trailing, shrubby and perennial, often several feet long, armed 
with numerous or very few prickles. Branches erect, 4 to 12 inches long, 
slightly pubescent, often prickly and glandular. Leaves three to seven- 
foliate, usually three-foliate; leaflets ovate or ovate-lanceolate, thin, pointed 
at the apex, rounded or narrowed at the base, sharply toothed. Flowers 
few in terminal racemes, white, about 1 inch broad; the five petals usually 
as long or slightly longer than the calyx lobes. Fruit black, usually at 
least one-half of an inch long and sometimes 1 inch long, fine flavored but 
with large seeds. 

In dry soil, especially in sandy sections, Nova Scotia to Ontario and 
Michigan, south to Virginia, Louisiana and Oklahoma. Flowering in May 
and June. Fruit ripe in June and July. 

A similar species, Rubus hispidus Linnaeus (Hispid or Run- 
ning Swamp Blackberry), with the stems densely beset with weak, retrorse 


bristles, 1s also common in swamps and low grounds throughout our area. 


Dalibarda; Dewdrop or False Violet 


Dalibarda repens Linnaeus 


Plate 97b 

Stems very slender, much tufted, several inches long. Leaves downy- 
pubescent on both sides, three-fourths to 2 inches broad, long petioled, 
ovate-orbicular, cordate, the margin with low, blunt or sometimes mucronu- 
late crenations; stipules setaceous. The scapelike peduncles 1 to 5 inches 
long, bearing one or two large, perfect white flowers, each flower about one- 
half of an inch broad; calyx deeply five or six-parted, three of the divisions 
larger than the other two or three; petals five; stamens numerous. Fruit 
composed of five to ten nearly dry drupelets, inclosed by the calyx segments. 
Stems also have short, recurved peduncles bearing several or numerous 
small cleistogamous flowers 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 100 


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WILD FLOWERS OF NEW YORK 139 


In rich woods, Nova Scotia to Minnesota, south to Pennsylvania, 
southern New Jersey, North Carolina, Ohio and Michigan. Flowering 
from June to September. 

Low or Pasture Rose 
Rosa virginiana Miller 
Plate 102 

A bushy shrub, a few inches to 3 or 4 feet high, sometimes higher; the 
stems armed with slender or stout, straight or curved infrastipular spines, 
and more or less prickly. Stipules entire. Leaves alternate with five or 
sometimes seven rather thin ovate-oval or obovate leaflets, dull green or 
somewhat shiny, coarsely toothed, one-half to 2 inches long, usually pointed 
at the end, glabrous or pubescent beneath. Flowers few or solitary, 2 to 3 
inches broad; pedicels and calyx usually glandular; calyx five-lobed, the 
segments lanceolate, long pointed, sometimes dilated toward the end, 
spreading and deciduous; petals five, obcordate, rose or pink, fading after 
opening. Fruit globose or depressed-globose, glandular-hispid, about one- 
third of an inch high. 

In dry or rocky soil, Newfoundland to Ontario and Wisconsin, south to 
Georgia, Louisiana and Missouri. Flowering from May to July. The 
Swamp Wild Rose (Rosa carolina Linnaeus) is frequent in open or 
wooded swamps and marshes. 


Apple Family 
Malaceae 
Black Chokeberry 
Aronia melanocarpa (Michaux) Britton 
Plate 103 

An extensively branching shrub, 3 to 8 feet high. Leaves obovate to 
oval, the apex varying from blunt to pointed, narrowed or cuneate at the 
base, short petioled, the margins crenulate, dark green above, paler beneath, 
glabrous or nearly so when mature. Flowers numerous in terminal, com- 
pound, leafy cymes; each flower about one-half of an inch broad; calyx and 


140 NEW YORK STATE MUSEUM 


pedicels glabrous or nearly so; calyx lobes glandular; petals five, concave, 
white or tinged with pink, spreading; stamens numerous, filaments rose- 
colored to white; anthers black or dark red. Fruit globose or oval, nearly 
black, or purplish black, about one-quarter of an inch in diameter. 

In low grounds, swamps or open woods, sometimes in drier situations, 
Nova Scotia to western Ontario, south to Florida and Michigan. Flowers 
in April and May. Fruit ripe in August and September. 

The Red Chokeberry (A. arbutifolia (Linnaeus) Elliott) has the 
cymes and lower surface of the leaves woolly and the mature fruit is bright 
red. The Purple-fruited Chokeberry (A. atropurpurea Britton) also 
has the cymes and lower leaf surfaces woolly but the mature fruit is 
purple-black. 

Senna Family 
Caesalpiniaceae 
Wild or American Senna 

Cassia marilandica Linnaeus 
Plate 104 

Stems 3 to.7 feet high, sparingly branched or simple, glabrous or with 
scant pubescence, from a perennial root. Leaves evenly pinnate, not 
sensitive to the touch, petioled and with a club-shaped gland near the base 
of the petiole; leaflets twelve to twenty, oblong, blunt but mucronate at the 
apex, rounded at the base, ciliate, 1 to 2 inches long, one-fourth to two- 
thirds of an inch wide. Flowers yellow, about two-thirds to three-fourths 
of an inch broad, numerous, in pubescent axillary racemes on the upper 
part of the plant. Calyx lobes five, nearly equal, ovate or oblong, obtuse; 
corolla nearly regular, of five spreading, nearly equal, clawed petals; 
stamens ten, the upper three imperfect. Fruit a flat linear pod, 3 to 4 
inches long and about one-fourth of an inch wide, curved, pubescent, con- 
taining flat, suborbicular seeds. The segments of the pod are about as 
long as broad. 

In moist meadows, marshes and swamps, sometimes on springy hill- 
sides, Massachusetts to central New York, Ohio, Tennessee and North 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum : ; Plate 102 — 


> 
| 
| 


LOW OR PASTURE ROSE 
Rosa virginiana 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 103 


BLACK CHOKEBERRY 
Aronia melanocar pa 


af 
i 


WILD FLOWERS OF NEW YORK I4I 


Carolina. Rather rare and local in New York, frequently seen along the 
Hudson River valley and up the Mohawk, northward along West Canada 
creek to Newport in Herkimer county, which appears to be the northern 
limit of its range. Flowering in July and August. 


Sensitive Pea; Wild Sensitive Plant 
Chamaecrista nictitans (Linnaeus) Moenchhausen 
Plate 105a 

An annual plant, 5 to 16 inches high with erect or decumbent, branching 
and somewhat pubescent stems. Leaves evenly pinnate, sensitive to the 
touch, bearing a small gland near the base of the petiole; leaflets twelve to 
forty, linear-oblong, blunt and mucronate at the apex, rounded and oblique 
at the base, inequilateral, one-fourth to two-thirds of an inch long, about 
one-fourth as wide. Flowers two or three together in the axils, one-fourth 
of an inch broad or less; calyx lobes five, pointed; corolla yellow, some- 
what irregular, three of the five petals smaller than the others; stamens 
five, all perfect. Fruit a small, linear, pubescent or smooth pod, 1 to 14 
inches long. 

In dry and sandy soil, Maine to Georgia, west to Indiana, Kansas 
and Texas. Flowering from July to September. 


Partridge Pea; Large-flowered Sensitive Pea 
Chamaecrista fasciculata (Michaux) Greene 
Plate 1o5b 

Stems annual, rather widely branched and pubescent with spreading 
hairs or nearly smooth, 1 to 2 feet high. Leaves with a sessile gland on 
the petiole, sensitive, similar to the preceding species but the twenty to 
thirty leaflets somewhat larger. Flowers two to four together in the axils, 
I to 14 inches broad and slender-pediceled; calyx lobes long pointed; petals 
yellow, sometimes purple spotted; stamens ten, all perfect; four of the 
anthers yellow, six of them purple. Fruit a linear, pubescent, or glabrous 
flattened pod, 13 to 23 inches long and one-fourth of an inch wide or less. 


142 NEW YORK STATE MUSEUM 


In dry or sandy soil, Massachusetts to Florida, Minnesota, Texas and 
Mexico. Flowering from July to September. 


Pea Family 
Fabaceae 
Wild Indigo; Horsefly Weed 


Baptisia tinctoria (Linnaeus) R. Brown 


Plate 1o7a 

Stems glabrous, erect, much branched, 2 to 4 feet high from a perennial 
root, blackening in dying. Leaves petioled, three-foliate, alternate; leaflets 
obovate, one-half to 13 inches long, nearly sessile, blunt, tapering at the 
base, entire. Flowers bright yellow, rather showy, in numerous, few- 
flowered, terminal racemes. Each flower about one-half of an inch long; 
calyx campanulate, the upper two lobes united into a lobe larger than the 
other three; corolla consisting of a standard (upper petal), two wings (lat- 
eral petals), and a keel (two lower petals); stamens ten, distinct; fruit a 
short ovoid or nearly globose pod, one-fourth to one-half of an inch long, 
and tipped with the subulate style. 

In dry or sandy soil, Maine to Vermont, Ontario, Minnesota, Florida 
and Louisiana. Flowering from June to September. 

The Blue Wild or Blue False Indigo, Baptisia australis 
(Linnaeus) R. Brown, has indigo blue flowers nearly an inch long and is 
naturalized in eastern and southern New York from the south. 


Wild or Perennial Lupine 


Lupinus perennis Linnaeus 
Figure XIX and Plate 106 


Stems 8 to 24 inches high, erect, pubescent and often branched, from 
a perennial root. Leaves digitately compound with seven to eleven (usually 
about eight), oblanceolate, sessile leaflets, blunt and mucronate at the 
apex, tapering to the base, I to 2 inches long, one-fourth to one-half of an 
inch wide, appressed-pubescent or glabrate; flowers blue, rarely white 
or pink, in terminal racemes; each flower one-half to two-thirds of an inch 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 


Plate 106 


WILD OR PERENNIAL LUPINE 
Lupinus perennis 


WILD FLOWERS OF NEW YORK 143 


long, on pedicels one-fourth to one-half of an inch long; calyx two-lipped; 
standard (upper petal) orbicular with reflexed margins, wings (two lateral 
petals) obovate; stamens monadelphous, with two forms of anthers. Fruit 
a linear-oblong, very hairy pod, 13 to 2 inches long and about one- 
fourth of an inch wide, with three to six seeds; the two valves of the pod 
coiling when it dehisces. 

In dry, sandy soil, Maine and Ontario to Minnesota, Florida, Missouri 
and Louisiana. Flowering in May and June. This is the only native 
species of Lupine in New York, although there are about fifty species in 


the western states. 


Figure XIX 
Wild or Perennial Lupine 


(Lupinus perennis Linnaeus) 


144 NEW YORK STATE MUSEUM 


Rabbit-foot, Old Field, or Stone Clover 
Trifolium arvense Linnaeus 
Plate ro8a 

An erect, usually much-branched annual, 5 to 18 inches high, silky- 
pubescent. Leaves very short petioled, three-foliate, oblanceolate or 
linear, minutely toothed, blunt and sometimes notched at the apex, nar- 
rowed at the base, one-half to 1 inch long. Flowers sessile in dense, ter- 
minal, peduncled, oblong or cylindric heads, one-half to 1 inch long; calyx 
very silky; corolla whitish, shorter than the elongated, slender, plumose 
calyx lobes. Fruiting pod very small. 

In waste places, dry and sandy fields, roadsides etc., Quebec and 
Ontario to South Carolina, Florida, Tennessee and Missouri. Naturalized 
from Europe. Flowering from May to September. 

The only Clover native to New York is the Buffalo Clover 
(Trifolium reflexum Linnaeus), which occurs from the western 
part of the State westward and southward. Our flora contains a large 
number of introduced species of clovers and related plants. Among them 
are the following: 


Vellowgomllop: Cloverime aces se eit ere ae rea ton eur! agrarium 
owxorpomallemilop: Clovers-osass46 one eer eer eee vy procumbens 
@rimsome Glovers certs to ae ais ected es See laren ean ies : incarnatum 
IRedkore VierdowsGloverians. seca eens neem ner pratense 
INIGIS Oe NkBoEIN (HOS aodaamecoon con sons aboss ene Ss hybridum 
AiWanttn(es ©) oniescs penne cco caieiere rues ai Oi Gere ciere oO bane 2 repens 
Alfalfa... :- ELM seicogn steed tee ce etayes tier SLL eco Eee ar EV Ie CLLR CLANCLO sativa 
iBlackwonsllopmVicdicae eer ena e ec ee . lupulina 
WihiterS weet" lovers wmecs marae s aetene ornate ee Melilotus alba 
WellowsS weet! Cloverimcnn «a vrcsat wre oe Cee € officinalis 


Bircksstootwlnetoil ments wot pices ocr e eee ete ee Lotus corniculatus 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 107 


+ 


A. WILD INDIGO; HORSEFLY WEED 
Baptisia tinctoria 


B. WHITE OR TRUE WOOD SORREL: ALLELUIA 
Oxalis acetosella 


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Goat’s-rue; Wild Sweet Pea 
Cracca virginiana Linnaeus 
Plate 1o8b 
Stems from a few inches to nearly 2 feet high, few or many in a dense 
cluster, erect or nearly so, from a stout, perennial root which is elongated, 
tough and fibrous. Stems and leaves silky with whitish hairs. Leaves 
odd-pinnate, short petioled; leaflets nine to twenty-five, oblong, linear- 
oblong or the terminal one oblanceolate, narrowed at the base, rounded and 
mucronate at the apex or sometimes notched, three-fourths to 1 inch long 
and one-eighth to one-third of an inch wide. Flowers crowded in a ter- 
minal, often compound and nearly sessile racemelike cluster; each flower 
one-half to three-fourths of an inch long on a short pedicel. Calyx with 
five nearly equal teeth; petals clawed, the standard rounded, yellow and 
conspicuous, wings and keel reddish or purplish. Fruiting pod narrow, 
densely hairy, 1 to 2 inches long. 
In dry and sandy soil, Maine to Minnesota, Arkansas, Florida, Louisi- 
ana and Northern Mexico. Flowering in June and July or sometimes as 
late as August in the north. 


Coronilla; Axseed; Axwort 


Coronilla varia Linnaeus 


Plate 109 
Stems ascending or straggling, glabrous and usually much branched, 
1 to 3 feet long from perennial roots. Leaves sessile; odd-pinnate; leaflets 
eleven to twenty-five, oblong or obovate, blunt and mucronate at the apex, 
narrowed or rounded at the base, one-half to three-fourths of an inch long, 
one-eighth to one-fourth of an inch wide. Flowers numerous in dense 
umbels terminating peduncles several inches in length; each flower one- 
third to one-half of an inch long on very short pedicels; standard (upper 
petal) pink, wings (lateral petals) white or purple-tipped. Fruit pod 
coriaceous, linear, four-angled, with two or three joints, each about one- 
fourth of an inch long or slightly longer. 


146 NEW YORK STATE MUSEUM 


Waste places, fields and roadsides, Massachusetts to southern New 
York, Maryland and Missouri. Adventive or naturalized from Europe. 
Flowering from June to August. 


THE TICK TREFOILS 
Meibomia 


The Tick Trefoils are all perennial herbs, often with stout roots, erect, 
ascending or trailing stems and three-foliolate leaves. The flowers are 
usually rather small, purplish in terminal or axillary, compound or simple 
racemes or panicles. Calyx two-lipped, the upper lip two-toothed, the 
lower lip with three acute or attenuate teeth. Stamens monadelphous 
or diadelphous (nine and one); anthers all alike. Fruit (loment) flat, sessile 
or stalked with several joints which are easily separable at maturity. 

A large genus of plants with about sixteen representatives in New 
York State. The following key is given as an aid in placing the various 
species not fully described and illustrated here. 


Loment not constricted above, deeply constricted below, long-stalked; leaflets broad 
Panicle terminal on the leafy stem 
Leaves crowded at the base of the panicle................ M. grandiflora 
eawes'scattered along-the stem ye. aa. sees sera ee ne M. pauciflora 
Panicle arising from the base of the plant, its stalk usually leatlesss 47.5 qaecee ee: 
M. nudiflora 
Loment constricted on both margins, more deeply below than above 
Stems trailing or reclining 
Leaflets orbicular, 1 to 2 inches long and pubescent......... M. michauxii 
Meafletstovate or oval: dulligreen’.: 4-5-2 soe ace eae ee ele lvamoreylalia) 
Stems erect or ascending 
Leaves sessile or nearly so; leaflets linear or lanceolate.....M. sessilifolia 
Leaves petioled 
Joints of the loment notably longer than broad 
Leaflets obtuse, yellowish green, rough-pubescent....M. canescens 
Weafletsvlong-acuminate.. 24ers eee eerie M. bracteosa 
Joints of the loment little longer than broad 


Loment distinctly long-stalked in the calyx 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 109 


CORONILLA; AXWORT: AXSEED 
Coronilla varia 


WILD FLOWERS OF NEW YORK 147 


Plants glabrous or nearly so (except varieties of M. paniculata) 
Leaflets lanceolate or oblong.............M. paniculata 
Leaflets broadly ovate or oval, glaucous beneath............. 

M. laevigata 

Plants pubescent 

Leaves velvety-pubescent beneath, thick and coriaceous....... 
MEE Wea ae sl Yel at at Il Ose ay 
Leaves appressed-pubescent or villous beneath and scarcely 
CORIACEOUS) ce a. Cee nr eee M. dillenii 

Loment sessile in the calyx or nearly so 
Loment joints four to seven, flowers showy; leaflets not coriaceous. . 
M. canadensis 

Loment joints one to three 
Leaflets scabrous, 1 to 2 inches long............M. rigida 
Leaflets not scabrous, one-half to 1 inch long 

Plant nearly glabrous throughout..M. marylandica 
Stem pubescent; leaflets and petioles ciliate.M. obtusa 


Prostrate Tick Trefoil 
Meibomia michauxti Vail 
Plate 110 
Stems prostrate, 2 to several feet long, softly pubescent or villous. 
Leaves petioled; leaflets nearly orbicular, pubescent, 1 to 2 inches long; 
stipules triangular-ovate, persistent, pointed, striate. Flowers in loose 
terminal and axillary panicles, purple, one-fourth to one-half of an inch 
long; calyx lobes ciliated. Loment about 1 inch long, three to five-jointed, 
the joints obliquely rhomboid below, slightly concave above, pubescent 
with hooked hairs, on a stipe about equaling the calyx lobes or slightly 
longer. 
Dry or sandy woods, New England and Ontario to Minnesota, Florida, 
Missouri and Louisiana. Flowering from July to September. 


148 NEW YORK STATE MUSEUM 


Large-bracted Tick Trefoil 


Meibomia bracteosa (Michaux) Kuntze 


Plate 111 

Stems erect, 2 to 6 feet high, glabrous or pubescent below, finely 
pubescent above in the panicle. Leaflets 2 to 8 inches long, longer than 
the petioles, ovate or ovate-lanceolate, long pointed, smooth or nearly so 
above, usually pubescent beneath; stipules lanceolate, sharp pointed. 
Flowers large, purple, one-third to one-half of an inch long with cuspidate, 
striate, deciduous bracts; calyx deeply two-lipped, the upper lip two-toothed. 
Loment 1 to 3 inches long, three to seven-jointed, the joints obliquely 
oblong, about twice as long as wide, pubescent with hooked hairs, on a 
stipe about as long as the lower calyx lobes. 

In open woods and thickets, Maine to Ontario, Minnesota, Florida, 
Missouri, Arkansas and Texas. Flowering in August and September. 


Dillen’s Tick Trefoil 
Meibomia dillenti (Darlington) Kuntze 


Plate 112 

Stems erect, pubescent with scattered hairs, 2 to 3 feet high. Leaflets 
rather thin, oval or oblong-ovate, blunt, 13 to 4 inches long, one-half to 
13 inches wide, sparingly pubescent or glabrous above, softly pubescent 
beneath; petioles much shorter than the leaflets. Flowers one-fourth to 
one-third of an inch long, purple, in loose, terminal, compound racemes 
with small, deciduous bracts. Loment 1 to 2 inches long, two to four- 
jointed, the joints nearly triangular, about one-fourth of an inch long, 
slightly convex on the back, pubescent with hooked hairs, with a stipe 
shorter than the calyx lobes. 

In woods and thickets, Maine to Ontario, Minnesota, Alabama, 
Tennessee, Missouri and Texas. Flowering from the last of June to 
September. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 7 — se = Plate 110 


PROSTRATE TICK TREFOIL 
Metbomia michauxti 


7 7 aan TT )6—| h(a eS ss 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum f Plate 111 


LARGE-BRACTED TICK TREFOIL 
Meibomia bracteosa 


i 


oa - et ieee 


er 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum 


Plate 112 


DILLEN’S TICK TREFOIL 
Meibomia dilleni 


WILD FLOWERS OF NEW YORK 149 


THE BUSH CLOVERS 
Lespedeza 


Herbs, often with perennial roots and erect or ascending stems. In 
a few species the stems trailing. Leaves three-foliolate. Flowers small, 
purple or whitish, in axillary clusters, heads or panicles. Frequently 
the flowers are of two kinds intermixed, one petaliferous but sterile, the 
other minute, apetalous and fertile. Calyx lobes nearly equal, those of 
the petaliferous flowers usually longer than those of the apetalous flowers. 
Upper petal (standard) obovate or oblong, the lower petals forming an 
incurved keel. Stamens usually diadelphous (nine and one); anthers all 
alike. Ovary one-ovuled, ripening into a flat, indehiscent, reticulated, 
mostly one-jointed and one-seeded pod. 


Key to the New York Species of Lespedeza 
Corolla purple or purplish; plants bearing both petaliferous and apetalous flowers 
Stems prostrate or trailing 
Foliage glabrous or somewhat appressed-pubescent 
Foliage downy-pubescent or tomentose 
Stems erect or ascending 
Peduncles distinct and mostly longer than the leaves 
Bushy-branched; petaliferous flowers paniculate...........L. violacea 
Stems simple or little branched; flowers racemose or subspicate 
Stems tomentose; leaves tomentose beneath........... WW, Jove 1 45 ©) hala st 
Stems and leaves glabrate or appressed-pubescent; leaflets oval to 
PPR cts ror ee clots ain a ps ceieic a Bore Dy a Gal lies 
Peduncles shorter than the leaves or the flower clusters sessile 


suborbicular 


Leaflets densely tomentose beneath; calyx of the petaliferous flowers less 

thankone-halivasslonguaskiac) POC === aia ee L. stuvei 
Leaflets appressed pubescent beneath or glabrate 

Calyx of petaliferous flowers less than one-half as long as the pod; 

leanletsyovaletorobloncc se ee eee L. frutescens 

Calyx of the petaliferous flowers two-thirds as long as the pod or more; 


leaflets silvery-pubescent beneath.................. L. simulata 


150 NEW YORK STATE MUSEUM 


Corolla whitish or yellowish; flowers all complete; pod included or scarcely exserted from 
the calyx 
Leaflets oblong, ovate-oblong, or nearly orbicular 
Pedunclesimostly exceeding tthetleaycs eon ee ert eee nr ree Lehi al 


Redunclesishonterthansthesyleavesmer pre er ie iaecieeeeee ee ESrcray puktayiia 


Stuve’s Bush Clover 
Lespedeza stuvei Nuttall 
Plate 113b 

Stems erect or nearly so, simple and wandlike or slightly branched, 
densely velvety or downy-pubescent all over, 2 to 4 feet high; petioles 
as long as the leaves or usually shorter; leaflets oval, oblong or suborbicular, 
blunt or retuse at the apex, one-half to 1 inch long. Flowers of both kinds 
in nearly sessile, axillary clusters; corolla violet-purple, one-fourth of an 
inch long or less. Pod oblong-ovate to orbicular, pointed, about one-fourth 
of an inch long or less, downy-pubescent, much longer than the calyx. 

In dry or sandy soil, Vermont and Massachusetts to Virginia, Alabama, 
Michigan, Arkansas and Texas. Flowering in August and September. 


Wandlike Bush Clover 
Lespedeza frutescens (Linnaeus) Britton 
Plate 114 

Stems erect, 1 to 3 feet high, simple or somewhat branched, finely 
pubescent or nearly smooth; petioles as long as the leaflets or shorter; 
leaflets oval, oblong or elliptic, blunt, truncate or notched at the apex, 
narrowed or rounded at the base, one-half to 13 inches long, smooth and 
dark green above, paler and pubescent beneath. Flowers of both kinds 
in short-stalked axillary clusters which are more or less crowded toward 
the summit of the stem; corolla violet-purple, about one-fourth of an inch 
long; pod ovate, pointed, pubescent, about one-fifth of an inch long. 

Dry soil in open woods, old fields, etc., Maine to Ontario, Minnesota, 
Florida, Hlinois and Texas. Flowering in August and September. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Z e Plate Bk 


A. HAIRY BUSH CLOVER B. STUVE’S BUSH CLOVER 
Lespedeza hirta Lespedeza stuvet 


7 


S 


WILD FLOWERS OF NEW 


é Memoir 15 N. Y. State Museum 


YORK 


Plate 114 


7 


WANDLIKE BUSH CLOVER 
Lespedeza frutescens 


WILD FLOWERS OF NEW YORK Si 


Hairy Bush Clover 
Lespedeza hirta (Linnaeus) Hornemann 
Plate 113a 

Stems rather stiff, erect and usually stout, simple or branched above, 
densely hairy or softly pubescent, 2 to 5 feet high. Leaflets three, oval 
or suborbicular, blunt at each end, or often notched at the apex, one-half 
to 2 inches long; the petioles shorter than the leaflets. Flowers in oblong- 
cylindric, rather dense heads on stalks which are much longer than the 
leaves; flowers all complete; corolla yellowish white, usually the standard 
with a purple spot near its base. Fruiting pod oval, pointed, hairy and 
about as long as the slender calyx lobes. 

In dry or sandy soil, Maine to Ontario and Minnesota, south to 
Florida, Louisiana and Texas. Flowering from August to October. 

The Round-headed Bush Clover (Lespedeza capitata 
Michaux) is similar but the stalks bearing the flower clusters are shorter 
than the leaves, forming a more compact inflorescence. Its range and 
period of flowering are about the same. 


Beach Pea; Seaside Pea 
Lathyrus maritimus (Linnaeus) Bigelow 

Plate 115 
Root perennial, stem glabrous or nearly so, stout, somewhat fleshy 
and slightly glaucous, angled, decumbent, 1 to 2 feet long with broad 
foliaceous, hastate, pointed stipules 1 to 2 inches long. Leaves nearly 
sessile, pinnate, the rachis terminating in a slender, forked tendril; leaflets 
six to twelve, oblong, oval or obovate, blunt and mucronulate at the 
apex, usually narrowed at the base, I to 23 inches long, one-half to 
three-fourths of an inch wide. Flowers six to ten on peduncles 3 to 4 
inches long, purple, three-fourths to 1 inch long; calyx teeth often ciliate. 
Fruit a sessile, linear-oblong, nearly glabrous, veined pod, 13 to 3 inches 

long and about one-half of an inch wide. 


152 NEW YORK STATE MUSEUM 


Sea beaches and sandy fields near the coast, New Jersey to Arctic 
America, also Oneida lake, Great Lakes, Pacific coast and in northern 
Europe and Asia. Flowering from May to August. 


Myrtle-leaved Marsh Pea 


Lathyrus myrtifolius Muhlenberg 


Plate 116 

Stems very slender, smooth, angled but not winged, weak, 1 to 3 feet 
long with obliquely ovate or half-sagittate stipules, one-half to 1 inch 
long, one-third to one-half of an inch wide and toothed; leaflets usually 
six, varying from four to eight, oval or ovate, pointed and mucronate at 
the apex, narrowed at the base, three-fourths to 2 inches long, one-fourth 
to two-thirds of an inch wide, the rachis terminating in a forked tendril. 
Flowering peduncles as long as the leaves or shorter, with three to nine 
flowers, yellowish in bud but turning purple as the flower opens; each 
flower about one-half of an inch long. Fruit a narrow, smooth pod without 
visible stalk, 1 to 2 inches long and somewhat less than one-fourth of an 
inch wide. 

In moist thickets, wet ground, swamps and shores, New Brunswick 
to Manitoba, south to North Carolina and Tennessee. Flowering from 
late in May until June or July. 

The members of the genus Lathyrus are often called Vetchlings, because 
of their close relationship to the true Vetches (genus Vicia), most of which 
are cultivated or naturalized species in our State. Another native Vetch- 
ling is the Marsh Vetchling (Lathyrus palustris Linnaeus), a 
boreal species found in the northern part of the State. It has linear leaflets 
and the stems are usually winged. The flowers are purple. The Cream- 
colored Vetchling (Lathyrus ochroleucus Hooker) with rather 
large cream-colored flowers, and broadly oval, acute leaflets, occurs 
throughout the western part of the State. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 115 » 


r 


a *¥ 


BEACH PEA; SEASIDE PEA 
Lathyrus maritinus 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 116 


MYRTLE-LEAVED MARSH PEA 
Lathyrus myrtifolins 


WILD FLOWERS OF NEW YORK 153 


Groundnut; Wild Bean 
Glycine apios Linnaeus 
Plate 117 
Stems slender, hairy or nearly smooth, with milky juice, climbing 
over herbs and bushes to a height of several feet, from a perennial root- 
stock of several necklace-shaped, edible tubers. Leaves pinnately com- 
pound, five- to seven-foliolate; leaflets ovate to ovate-lanceolate, pointed 
at the apex, rounded at the base, 1 to 3 inches long. Flowers brownish 
purple, fragrant, about one-half of an inch long, in axillary racemes; 
peduncles shorter than the leaves; rachis of the inflorescence knobby; 
calyx two-lipped, the two lateral teeth very small, the two upper united 
and short, the lower one long and acute; standard ovate or orbicular and 
reflexed, wings obliquely obovate, adherent to the elongated, incurved 
and at length twisted keel; pod narrow, straight or slightly curved, 2 to 
43 inches long and about one-fourth of an inch wide or less, many-seeded 
and rather thick in texture. 
Moist thickets along streams, bottomlands, or low woods. New 
Brunswick to Florida, west to Ontario, Minnesota, Nebraska, Kansas and 
Texas. Flowering from July to September. 


Wild or Hog Peanut 
Falcata comosa (Linnaeus) Kuntze 
Plate -118a 
Stems very slender, simple or somewhat branched, twining and 
climbing over herbs and shrubs, 1 to 6 feet long, more or less pubescent. 
Leaves with three rhombic-ovate or broadly ovate leaflets pointed at the 
apex, rounded at the base, 1 to 3 inches long. Flowers purplish or nearly 
white in axillary, slender-stalked clusters or racemes. In the lower axils 
are solitary, apetalous, fertile flowers. Calyx of the petaliferous flowers 
four to five-toothed, tubular; the oblong wings of the corolla curved 
and adherent to the recurved, blunt keel and inclosed by the erect, 


154 NEW YORK STATE MUSEUM 


obovate standard; pods oblong-lanceolate, pointed and hairy, about 1 
inch long. 

In moist, shaded places, New Brunswick to Florida, west to Manitoba, 
Nebraska and Louisiana. Flowering from early in August until late in 
September. 

Falcata pitcheri (Torrey, & Gray) Kuntze, very closely 
related to the preceding species, has leaves of a firmer texture and the 
stems, petioles and flowering stalks villous-pubescent with conspicuous 
brown hairs. 

Trailing Wild Bean 
Strophostyles helvola (Linnaeus) Britton 
Plate 118b 

A twining or trailing and climbing, herbaceous, rough-pubescent vine. 
Stems more or less branched below, 2 to 7 feet long, or dwarfed and almost 
erect, from an annual root. Leaves pinnately three-foliolate; leaflets ovate, 
pointed or blunt at the apex, the base rounded, thickish in texture, usually 
bluntly lobed, 1 to 2 inches long, the two lower leaves unequal at the base. 
Flowers greenish purple, about one-half of an inch long, three to twelve 
together in dense, capitate clusters at the ends of long, axillary stalks which 
are longer than the leaves; keel of the corolla strongly curved and slender. 
Fruiting pod round in cross-section, somewhat hairy, linear and without a 
stalk, 13 to 3 inches long. 

In sandy fields and thickets, mainly near the coast, Quebec to Massa- 
chusetts and Florida, less frequent westward through Ontario to South 
Dakota and Kansas. Flowering from the latter part of July to September 
and October. 

A closely related species of Long Island and southward, S. umbel- 
lata (Muhlenberg) Britton, has shorter pods, slightly larger flowers, 
entire leaflets and perennial roots. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 117 
i 7 


GROUNDNUT; WILD BEAN 
Glycine apios 


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MUOA MON AO SHUAMOTA GCIIM 


WILD FLOWERS OF NEW YORK 155 


Geranium Family 
Geraniaceae 
Herb Robert; Red Robin 
Robertiella robertiana (Linnaeus) Hanks 


(Geranium robertianum Linnaeus) 


Plate 119 

Roots mostly biennial, sometimes annual, giving rise to one or several 
ascending or nearly erect, glandular-pubescent stems 5 to 18 inches high. 
Entire plant with a strong, disagreeable odor. Leaves rounded-ovate in 
outline, the divisions deeply cleft or lobed, the margins with oblong, mucro- 
nate teeth. Flowers reddish purple, about one-half of an inch broad, two 
on each stalk; sepals five, each tipped with an awn. Petals five, each with 
a slender claw and an obovate, rounded blade. Stamens ten; ovary five- 
lobed and five-celled. Fruiting capsule about 1 inch long, awn-tipped, 
separating at maturity into five carpels, the bodies deciduous from the 
styles at maturity, each with two fibrous appendages near the top. 

In rich soil of rocky woodlands, Nova Scotia to Manitoba south to 
Pennsylvania and Missouri; also in Europe and Northern Africa. Flow- 
ering from May to September. 


Wild Geranium; Wild or Spotted Crane’s-bill 


Geranium maculatum Linnaeus 


Plate 120 

Stems mainly simple from a stout, perennial rootstock, often much 
branched above, hairy, 10 to 20 inches high. The basal leaves nearly 
orbicular, broadly heart-shaped, on long leaf-stalks; the blades 3 to 5 inches 
wide, deeply three to five-lobed with wedge-shaped divisions, the margins 
cleft or toothed; leaves of the stem two, opposite, short-stalked, similar 
to the basal leaves. Flowers rose-purple, 1 to 13 inches broad, terminal 
in two to five-flowered, loose, leafy-bracted umbels; sepals sharp pointed; 
petals five, woolly at the base, thin, broad and overlapping one another. 
Fruit an elongated capsule tipped with the persistent compound style, 


156 NEW YORK STATE MUSEUM 


which in fruit is I or more inches long; carpels of the fruit permanently 
attached to the styles, separating from the base and curved upward in 
dehiscence. 

In rich or moist woodlands, Maine and Ontario west to Manitoba, 
south to Georgia and Alabama. Flowering from late in April to June 
or July. 

There are several other species of Geranium in New York, most of 
them small-flowered, introduced species. Of the native ones, Bicknell’s 
Geranium (Geranium bicknellii Britton) is an annual with small, 
purple flowers, somewhat less than one-half of an inch broad in a loose 
cluster, on two-flowered peduncles, with the beak of the fruit long pointed. 
The Carolina Geranium (Geranium carolinianum Linnaeus) 
has a more compact inflorescence and a short-pointed fruit. 


Wood Sorrel Family 
Oma liidkaxciesare 
White or True Wood Sorrel; Alleluia 


Oxalis acetosella Linnaeus 


Plate 107b 

Rootstock perennial, slender, scaly and little branched. Leaves basal, 
three to eight together, each 2 to 6 inches high, pubescent; petioles broad- 
ened at the base and jointed. Leaflets three, obcordate, wider than long, 
one-half to 1 inch wide. Flowers one-half to three-fourths of an inch 
broad, solitary on stalks as long or longer than the leaves; petals white 
or pink with deep pink veins, three or four times longer than the 
calyx; stamens ten. Fruit a subglobose, cylindric capsule about one- 
sixth of an inch long. At the base of the leaves are also found short-stalked 
flowers. 

In cool, damp woods, or shaded mossy banks, Nova Scotia to the north 
shore of Lake Superior, south to North Carolina. Also in Europe, Asia 
and northern Africa. Flowering from May to July. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 119 


HERB ROBERT; RED ROBIN 
Robertiella robertiana 


i ; : ; i = 


cia ee 2 ae a 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 120 


\ 


WILD GERANIUM; WILD OR SPOTTED CRANE’S-BILL 
Geranium maculatum 


WILD FLOWERS OF NEW YORK 15 


N 


Violet Wood Sorrel 


Tonoxalis violacea (Linnaeus) Small 


Plate r2ta 

Flowering stalks and leaves smooth, 3 to 8 inches tall, arising from a 
perennial, brownish, scaly bulb. Leaves few or several, slender-stemmed, 
one-half to 1} inches wide. Leaflets three, broader than long, notched at 
the apex. Flowers three to ten, or rarely more on each stalk, forming a 
loose, umbellate inflorescence at the summit, which is taller than the leaves. 
Each flower two-thirds to three-fourths of an inch long on a short, slender 
pedicel; sepals blunt, five in number, with tubercles at the apex; petals five, 
rose-purple, lighter toward the base, blunt, about three times as long as the 
sepals; stamens ten; capsule ovoid, about one-fifth of an inch in diameter. 

In open, usually rather dry woodlands, shaded hillsides and thickets; 
sometimes in open, recently cleared land, Massachusetts to Florida and 
Texas, west to Minnesota. Flowering in May and June. 


Tall Yellow Wood Sorrel 


Xanthoxalis cymosa Small 


Plate r21b 

Stems ascending or erect, branched above, 6 inches to 3 or 4 feet high 
and frequently reclining on surrounding vegetation, usually hairy, reddish 
or brown. Leaves bright green, three-fourths to 1} inches broad on petioles 
1 to 3 inches long; leaflets broader than long, sharply notched. Flowers 
yellow, in forking cymes; each flower on a pedicel one-fourth to one-third 
of an inch long, which is more or less hairy and erect or ascending. Sepals 
five, lanceolate or narrowly elliptic, one-sixth to one-fourth of an inch long, 
spreading in fruit; petals five, obtuse or notched at the apex, one-third to 
nearly one-half of an inch long. Fruit a slender, columnar, erect capsule, 
about one-half of an inch long, gradually narrowed to the summit; seeds 
obovoid-oblong with nearly continuous ridges. 

In fields, thickets and woods, Ontario to Michigan, south to Florida 
and Texas. Flowering from May to October. 


158 NEW YORK STATE MUSEUM 


There are several closely related species of Yellow Sorrel. Those of 
the northeastern states are illustrated and described by Britton & Brown 
@ilitus. Flora, 2: 432-35; ed: 2) (XS sta ueita, 26 bits hid) Xe er aa 
and) x<«, bird ¢ toma e): 

Jewelweed Family 
Balsaminaceae 
Spotted or Wild Touch-me-not 
Impatiens biflora Walter 


Plate 122a 

A tall, glabrous annual, 2 to 6 feet high and much branched, more 
or less purplish. Leaves alternate, thin, ovate and elliptic, glaucous 
beneath, 1 to 3 inches long, blunt, the margins toothed. Flowers horizontal, 
orange-yellow, mottled with reddish brown, or rarely nearly white and 
not mottled, three-fourths to 1 inch long, on slender, pendant stalks. Sepals 
three, the two lateral ones small, green, nerved, the other one large, conic, 
petallike, saccate and spurred, longer than broad, contracted into a slender 
incurved spur, two-toothed at the apex. Petals three, with two of them 
two-cleft into dissimilar lobes; stamens five. Fruit an oblong capsule, 
violently and elastically dehiscent at maturity into five spirally coiled 
valves, expelling the oblong, ridged seeds. Also developing small, 
cleistogamous flowers later in the season. 

Low grounds, thickets, ditches, along streams and low, moist wood- 
lands, Newfoundland to Saskatchewan, south to Florida and Nebraska. 


Flowering from July to September. 


Pale Touch-me-not; Jewelweed 
Impatiens pallida Nuttall 


Plate 122b 
Resembling the Spotted Touch-me-not, but usually stouter and higher. 
Flowers pale yellow, sparingly spotted with reddish brown or without 
spots, 1 to 14 inches long, the saccate sepal dilated-conic, as broad as 
long, abruptly contracted into a short, scarcely incurved spur, which is 
less than one-third the length of the saccate sepal. 


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WILD FLOWERS OF NEW YORK 159 


In situations similar to the preceding species but more common north- 
ward, Nova Scotia to Saskatchewan, south to Georgia and Kansas. 
Flowering from July to September. 


Milkwort Family 
Polygalaceae 


A family of small herbs (our species) with alternate, opposite or 
whorled leaves. Flowers racemose, spicate or capitate, rarely solitary 
and axillary, sometimes also with cleistogamous and subterranean flowers. 
Sepals very unequal, the two lateral ones large and petallike. Petals 
three, united into a tube which is split on the back, and more or less adnate 
to the stamens. Stamens eight or six, monadelphous below, or diadelphous. 
Capsule membranaceous, compressed, dehiscent along the margin. Seeds 
one in each cavity of the capsule and usually hairy. 

Our species all belong to Polygala, a very large genus of plants, con- 
taining about sixty species in North America, of which about eleven are 
found in New York State. The following key may be of service in 
identifying them. 


Flowers orange-yellow in a dense oblong spike; basal leaves spatulate....... w 12 Itlii@e 
Blowers roselon punplew distinctly racemose pene eee eer 2P. polygama 
Flowers rose-purple to white, one to four in number, axillary, but apparently terminal... . 
Bes piaracn folira 
Flowers in terminal, more or less elongated spikes, or if the spikes oblong, flowers not 
yellow and no basal leaves 
Leaves at least the lower, verticillate, spikes 4 to g lines thick, blunt; flowers purple 
to greenish white 

Spikes sessile or nearly so; wings deltoid.................... A 12, CRB ONE 6S) 
Spikes peduncled; wings lanceolate-ovate................. 5 1% lorevirolia, 

Leaves verticillate and alternate; spikes 2 to 3 lines thick and acute 
Verticillate leaves predominating; spikes dense; flowers green to purplish........ 
Gl. werrnenilila we, 
Alternate leaves predominating; spikes loose; flowers more purple 


yj 12k, yxaa\ oy feriel ee 


160 NEW YORK STATE MUSEUM 


Leaves all alternate 
Petals united into a tube which is cleft and about one-fourth of an inch long; 


HOWwers) pinks. eee RT tet cccrer ne eer ene eee SePs anican nia tra 
Petals not conspicuously united into a tube 
Spikes ovoid to globose; bracts persistent; flowers rose-purple to white...... 
9 P. viridescens 
Spikes cylindric 
Leaves oblanceolate to linear, 2 to 6 lines long; flowers greenish to 
DUTP lish Macs wae ne Ae eee Pacis or oer TOPs niu tite lis. 


Leaves lanceolate, 1 to 2 inches long; flowers white to greenish........ 
Tr Ps slemierrca) 
Orange Milkwort; Wild Bachelor’s-button 
Polygala lutea Linnaeus 
Plate 123a 
Stems annual, smooth, tufted from fibrous roots, erect or ascending, 
sometimes becoming branched, 6 to 12 inches high. Stem leaves oblong- 
lanceolate, pointed or blunt, three-fourths to 14 inches long, one-fourth 
of an inch wide or less, entire; basal leaves broader and usually larger, 
obovate or spatulate, blunt. Flowers in terminal, blunt, spikelike racemes 
which are dense and ovoid or oblong in shape, one-half to 14 inches long, 
one-half to three-fourths of an inch thick. Individual flowers about one- 
fourth of an inch long, orange-yellow, preserving their color in drying; 
wings oblong-ovate, abruptly pointed; crest of the corolla tube minute; 
caruncle lobes linear, about equaling the hairy seed, or shorter. 
In pine-barren depressions and swamps, Long Island to New Jersey 
and eastern Pennsylvania to Florida and Louisiana. Flowering from 
June to August or September. 


Cross-leaved or Marsh Milkwort 


Polygala cruciata Linnaeus 


Plate 123b 
A small annual, 3 to 15 inches tall, with three or four alternate branches 


above, the stems angled or square. Leaves all on the stem and mostly 


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WILD FLOWERS OF NEW YORK 161 


verticillate in fours, oblanceolate or linear-lanceolate, one-half to 14 inches 
long and about one-eighth of an inch wide, entire, blunt and mucronulate 
at the apex. Flowers in short-stalked, oval, blunt racemes, about one- 
third to one-half of an inch thick, purplish green or nearly white; wings 
triangular-ovate, sessile, somewhat heart-shaped, pointed or awned, one- 
fourth of an inch long or less, longer than the pods; crest of the corolla 
minute; seeds oblong and somewhat hairy. 

In low grounds and marshes along the coast and sandy swamps inland, 
Maine to Florida west to Michigan, Minnesota, Nebraska and Louisiana. 
Flowering from July to September. 


Field or Purple Milkwort 


Polygala viridescens Linnaeus 
Plate r24a 

Stems leafy, branched above, annual, smooth and somewhat angled, 
5 to 15 inches high. Leaves all on the stem, oblong to linear-oblong 
in shape, three-fourths to 13 inches long, one-eighth or one-sixteenth of 
an inch wide, pointed and mucronulate. Flowers purplish, greenish purple, 
whitish or greenish in terminal, globose, blunt heads about one-half 
of an inch thick, becoming oval; wings of the flowers sessile, ovate, often 
slightly cordate, longer than the pod, bracts usually persistent on the 
elongating axis as the lower flowers fall away. 

In meadows, fields and sandy depressions, Nova Scotia to Ontario 
and Minnesota, south to North Carolina, Kansas and Arkansas. Flowering 
from June to September. 


Seneca Snakeroot; Mountain Flax 
Polygala senega Linnaeus 
Plate r25a 
Stems usually several from a perennial, woody rootstock, ascending 
or erect, 5 to 18 inches high, usually simple, occasionally branched above, 
glabrous or nearly so. Leaves alternate, the lowest ones very small, closer 
together and scalelike, the upper ones oblong-lanceolate or ovate, some- 


162 NEW YORK STATE MUSEUM 


times lanceolate, sessile, 1 to 2 inches long, about one-fourth to one-third 
of an inch wide, minutely toothed. Flowers white, sometimes tinged with 
green, in dense, terminal, pointed spikes, I to 2 inches long. Each flower 
about one-eighth of an inch long; wings of the flower orbicular-obovate, 
concave; crest of the corolla short and few-lobed. 

In dry or rocky woodlands, New Brunswick to Hudson bay and 
Alberta, south along the mountains to North Carolina and west to Missouri 
and Arkansas. Flowering in May and June. 


Racemed Milkwort 
Polygala polygama Walter 
Plate 124b 

Stems usually several or many froma deep, slender, perennial root, 
smooth and simple, 4 to 15 inches high. Leaves crowded along the stems, 
oblong or broadly lanceolate, blunt, mucronulate, two-thirds to 1 inch 
long and about one-sixth of an inch wide or less, the lower leaves usually 
smaller. Flowers purple, rarely whitish, in a loose, terminal raceme, 1 to 
3 inches long or less; wings of the flower broadly ovate; crest of the corolla 
large and fringed; stamens eight. Numerous cleistogamous flowers are 
developed upon conspicuous, whitish subterranean branches which rise from 
the base of the stems. 

In dry or sandy fields and meadows, Nova Scotia to Manitoba, south 
to Florida, Texas and Michigan. Flowering in June and July. 


Fringed Milkwort; Flowering Wintergreen 
Polygala paucifolia Willdenow 
Plate 125b 
Flowering and leaf-bearing stems smooth, ascending or erect, 3 to 7 
inches high from slender, prostrate, perennial stems and rootstocks which 
are often several inches in length. Leaves few, clustered at the summit of 
the stems, ovate or oblong, 1 to 14 inches long, two-thirds to 1 inch wide, 
pointed at each end, rough-margined. Flowers one to five on a_ stem, 


axillary to the upper leaves, rose-purple or rarely white, two-thirds to 


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WILD FLOWERS OF NEW YORK 163 


1 inch long on slender, short flower stalks; wings of the flower obovate; 
crest of the corolla beautifully fringed. The base of the stems and root- 
stocks bear numerous cleistogamous flowers on short, lateral, subterranean 
branches. 

In rich, moist woods, New Brunswick to Saskatchewan south to 
Georgia, Illinois and Minnesota. Flowering in May and June. 


Buckthorn Family 
Rhamnaceae 
New Jersey Tea; Redroot 
Ceanothus americanus Linnaeus 
Plate 126 

A low shrub with several or many ascending stems from a deep 
reddish root. Leaves alternate, ovate or oblong-ovate, I to 3 inches 
long, one-half to 14 inches wide, pointed at the apex, blunt or slightly 
heart-shaped at the base, pubescent, especially beneath, the margins finely 
toothed, the blade conspicuously three-nerved; petioles usually less than 
one-half of an inch long. Flowers small and white, in dense, oblong clusters 
on terminal or elongated axillary stalks; limb of the calyx tube five-lobed; 
petals five, with narrow claws and bearded blades; stamens five, their 
filaments slender and elongated. Fruit dark colored or nearly black when 
mature, three-lobed, about one-sixth of an inch long. 

In sandy or dry open woods and fields, Maine to Ontario and Manitoba 
south to Florida and Texas. Flowering from the latter part of May to 


July. 
Mallow Family 


Malvaceae 
Marsh Mallow; Wymote 


Althaea officinalis Lannaeus 


Plate 127 
An introduced plant, well established in certain salt marshes along the 
coast, with erect, herbaceous stems, 2 to 6 feet high from perennial roots, 


164 NEW YORK STATE MUSEUM 


densely velvety-pubescent all over. Leaves ovate, pointed or blunt at 
the apex, toothed and usually somewhat three-lobed, the lower leaves 
usually cordate, veins conspicuously raised on the lower surfaces. Flowers 
pink or nearly white, about 14 inches broad in terminal and axillary leafy 
clusters. Calyx segments five, ovate-lanceolate, subtended by six to nine 
linear bractlets; petals five; stamens numerous, forming a central column 
around the pistil and united with the bases of the petals; styles united 
below. Carpels fifteen to twenty, each one-seeded, arranged in a circle 
around the axis of the fruit. 

Flowering in June and July. The photograph for the illustration of 


this species was taken near Port Washington, Long Island. 


Musk Mallow; Musk Plant 


Malva moschata Linnaeus 
Plate 128 

Stems erect, I to 23 feet high, more or less branching and hairy, from 
a perennial root. Leaves orbicular in outline, 3 to 4 inches broad with 
several broad, rounded, toothed lobes; stem leaves deeply cut into narrow 
segments. Flowers 13 to 2 inches broad, pink or white, slightly musk- 
scented, clustered in leafy racemes at the summits of the stems and branches; 
petals five, notched at the apex, several times longer than the pointed, 
triangular-ovate calyx lobes; stamens numerous, forming a column in the 
center of the flower; carpels of the fruit fifteen to twenty in number, densely 
hairy, rounded at the back. 

Roadsides, fields and waste places throughout the eastern states. 
Native of Europe and thoroughly naturalized in many places. Flowering 
from July to September. 

Other Mallows, native of the Old World and adventive or natural- 
ized in the eastern states, are the High Mallow (Malva sylvestris 
Linnaeus), the Low, Dwarf or Running Mallow, also known as Cheeses 
(Malva rotundifolia Linnaeus), the Whorled Mallow or Curled 
Mallow (Malva verticillata Linnaeus) and the Vervain Mallow 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 126 


NEW JERSEY TEA; REDROOT 
Ceanothus americanus 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 127 


MARSH MALLOW; WYMOTE 
Althaea officinalis 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 128 
[ SS ee —— == ——e a = | 


MUSK MALLOW; MUSK PLANT 
Malva moschata 


WILD FLOWERS OF NEW YORK 165 


(Malva alcea Linnaeus). Descriptions of these may be found in the 
current floras or manuals of botany of the northeastern states. 


Swamp Rose Mallow; Mallow Rose 
Hibiscus moscheutos Linnaeus 
Plate 129 

Stems tall and canelike from a perennial root, 3 to 6 feet high. Leaves 
ovate-lanceolate or ovate, blunt or rounded and often slightly heart- 
shaped at the base, pointed or blunt at the apex, 3 to 5 inches long, the lower 
ones sometimes lobed, all conspicuously palmately veined, toothed, densely 
hairy with white stellate hairs beneath, green and finely hairy or nearly 
smooth above. Flowers 4 to 7 inches broad, pink, clustered at the top of 
the plant; calyx lobes five, ovate, pointed, subtended by several narrow 
bractlets; petals five, broadly obovate; stamens numerous in a column 
surrounding the style which is five-cleft at the summit with five stigmas. 
Fruit a five-chambered pod about 1 inch long, blunt or slightly pointed. 

In marshes along the ocean or near the coast from eastern Massachusetts 
to Florida and in saline situations and marshy lake shores inland, especially 
throughout the Great Lakes region. Flowering in August and September. 

The Crimson-eye Rose Mallow (Hibiscus oculiroseus Britton) 
is similar, but the flower is white with a dark-crimson center and the fruit 
pod is long pointed. It is found on Long Island, Staten Island and in New 


Jersey. 
Saint John’s-wort Family 
Hy pernicaceae 

A family containing about sixteen species in New York State, mostly 
herbs, some of them shrubs, chiefly with opposite leaves and yellow or 
rarely reddish flowers in terminal clusters. In many of them the foliage 
is pellucid-punctate or dotted. Flowers regular and perfect. Sepals 
four or five. Petals four or five. Stamens numerous or few, often in 
three or five sets. 


166 NEW YORK STATE MUSEUM 


Key to the New York State Species of the Saint John’s-wort Family 


Sepals four, in unequal pairs; petals also four 
Stems erect, 1 to 2 feet high; leaves clasping, styles three to four................... 
Tt AUSiChyen Wm) Sitasns 
Stems diffusely branched, 5 to ro inches high; leaves sessile; styles two............. 
2 Ascyrum hypericoides 
Sepals and petals usually five 
Petals pink or greenish purple, imbricated in the bud; leaves sessile; capsules red- 


[SIU OM See ae nic ree chy caw OS WIN oN oe Gato g Ceti d c 3 Triadenum virginicum 


Petals yellow, convolute in the bud 
Leaves reduced to minute appressed scales...4 Sarothra gentianoides 
Leaves normal 
Styles five, large perennials 
Flowers 1 to 2 inches broad; capsules three-fourths of an inch long..... 
5 Hypericum ascyron 
Flowers one-half to 1 inch broad; capsules about one-half of an inch 
LOM) tenet Pane ers tenet aeteae sta ieee 6 Hypericum kal mianulm 
Styles three or rarely four 
Tall leafy shrubs with numerous flowers 

Flowers one-half to three-fourths of an inch broad; pods one-third 
to one-half of an inch long...7 Hypericum prolificum 
Flowers one-third to one-half of an inch broad; pods one-sixth to 
one-fourth of an inch long...8 Hypericum densiflorum 

Herbaceous perennials, or woody at the base only 
Stamens numerous (fifteen to forty); flowers one-fourth to 1 inch 

broad 
Capsules one-celled or incompletely three to four-celled 
Capsules incompletely three to four-celled by the pro- 
jectine’ placentae-leaves oblonge.. 1-445 cn eee oe 
9 Hypericum adpressum 
Capsules strictly one-celled; placentae parietal; styles 
united into a beak, separate above; leaves elliptic...... 
io Hypericum ellipticum 
Capsules completely three-celled; styles separate 

Leaves linear or oblong; sepals lanceolate. Introduced 


from Europe.....11 Hypericum perforatum 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 129 


SWAMP ROSE MALLOW; MALLOW ROSE 
Fibiscus moscheutos 


—_ =e - 


Se 
> 


WILD FLOWERS OF NEW YORK 167 


Leaves broadly oblong, oval or ovate-lanceolate, native; 
sepals ovate to ovate-lanceolate, acute 


12 Hypericum punctatum 
Stamens few (five to twelve); flowers one-tenth to one-fourth of 
an inch broad 
Cyme leafy-bracted ......... 13 Hypericum boreale 
Cymes subulate-bracted 
Leaves ovate, oval or oblong; capsules one-twelfth to 


five-twelfths of an inch long 


14 Hypericum mutilum 
Leaves lanceolate or oblong-lanceolate; capsule one-third 


to one-half of an inch long 


I5 Hypericum majus 
Leaves linear, blunt, three-nmerved..................-:-- 


16 Hypericum canadense 


Great or Giant Saint John’s-wort 


Hypericum ascyron Linnaeus 


Plate 130a 

Stems herbaceous, 2 to 5 feet tall from a perennial root, angled, 
branching and smooth. Leaves sessile, ovate-oblong to ovate-lanceolate, 
2 to 5 inches long and three-fourths to 13 inches wide, clasping the stem. 
Flowers bright yellow, showy, I to 2 inches broad, few or several in a loose, 
terminal cluster. Sepals five, ovate-lanceolate, about one-half of an inch 
long, pointed; petals five, obovate or oblanceolate; stamens numerous, 
united into five sets. Styles usually five in number, united below, the 
stigmas capitate. Fruit pod ovoid in shape, three-fourths to seven-eighths 
of an inch long. 

Chiefly along streams, Quebec to Vermont and Manitoba south to 
Connecticut, Pennsylvania, Illinois and Missouri. Flowering in July 
and August. 

The St Peter’s-wort (Ascyrum stans Michaux) and the St 
Andrew’s Cross (Ascyrum hypericoides Linnaeus), two small, 
leafy, shrubby species of the coastal region, differ from the species of 


168 NEW YORK STATE MUSEUM 


Hypericum by having four instead of five petals, as pointed out above in 
the key to the species of the family. 


Elliptic-leaved or Pale Saint John’s-wort 
Hypericum ellipticum Hooker 
Plate 130b 

Stems herbaceous, slightly four-angled, mainly simple or with a few 
branches, erect, 6 to 20 inches high from a perennial root. Leaves oval 
or elliptic, sessile, spreading, thin in texture, three-fourths to 11 inches 
long and one-fourth to one-half of an inch wide, blunt at the apex. Flowers 
pale yellow, few or several in terminal cymes, each flower about one-half 
of an inch broad, central flower of each cluster opening first, the lateral 
branches of the cluster developing later; sepals slightly shorter than the 
petals; styles three, united below; capsules or fruit ovoid-globose, about 
one-eighth of an inch long and one-celled. Sepals and petals occasionally 
four instead of five. 

In low grounds, swamps and along streams, Nova Scotia to Manitoba, 
south to New Jersey, Maryland and Minnesota. Flowering in July and 
August. Our illustration was made from plants just beginning to flower, 
before the lateral branches of the cyme were developed. 


Common Saint John’s-wort 
Hypericum perforatum Linnaeus 
Plate 131a 

Stems herbaceous, smooth, slender, 1 to 2 feet high from a perennial 
root, usually much branched and with several barren shoots at the base. 
Leaves sessile, linear or oblong, one-half to 1 inch long, one-tenth to one- 
third of an inch wide, blunt, black dotted. Flowers three-fourths to 1 
inch broad, bright yellow, several or many in terminal cymes; petals five, 
black dotted, longer than the lanceolate pointed sepals; stamens numerous, 
united by their base into three sets; styles three; fruit pod or capsule ovoid, 

about one-fourth of an inch long or less, three-celled. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 130 


L 


A. GREAT OR GIANT SAINT JOHN’S-WORT B. ELLIPTIC-LEAVED OR PALE 
Hypericum ascyron SAINT JOHN’S-WORT 
Hypericum ellipticum 


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WILD FLOWERS OF NEW YORK 169 


Native of Europe and naturalized as a weed in fields and waste places 
throughout the East. 
Rockrose Family 
Cis trarcierare 
Frostweed; Rockrose 
Crocanthemum canadense (Linnaeus) Britton 
Plate 131b 

Stems erect or diffuse from a perennial root, 5 to 20 inches high, finely 
canescent and becoming branched. Leaves nearly sessile, linear-oblong to 
oblanceolate, one-half to 14 inches long, one-third of an inch wide or less, 
green above, canescent beneath. Flowers bright yellow, usually one or two, 
1 to 12 inches broad; sepals five, the two outer ones much smaller; petals 
five, broadly obovate, soon withering and falling; capsule ovoid, about one- 
third of an inch long. After the petaliferous flowers fade and fall, the 
axillary branches elongate and bear numerous apetalous sessile flowers, 
which develop fruiting capsules about one-sixth of an inch in diameter. 

In sandy fields or rocky soil, Maine to Ontario and Wisconsin, south 
to North Carolina and Mississippi. Flowering from May to July. 

A closely related species of similar situations, Crocanthemum 
majus (Linnaeus) Britton, is more canescent, the primary flowers 
clustered at the summit of the stem and not surpassed by the branches; 
the secondary or apetalous flowers very small and producing capsules 
which are only about one-twelfth of an inch in diameter. 


Woolly Hudsonia; False Heather 
Hudsonia tomentosa Nuttall 
Plate r32a 
A diffusely branched, low-tufted or matted perennial, somewhat 
woody, 3 to 8 inches high, pale and hoary-pubescent, the branches stout and 
ascending. Leaves small and scalelike, about one-tenth or one-twelfth of 
an inch long, overlapping one another and appressed to the stem. Flowers 
numerous, almost sessile or on short, stout stalks less than one-fourth of an 


170 NEW YORK STATE MUSEUM 


inch long, yellow, about one-fourth of an inch broad or slightly less; petals 
five, obovate-oblong; stamens numerous, nine to eighteen in number; 
sepals blunt. Fruit a small, ovoid, smooth and one-seeded capsule. 

In sandy pine barrens and sandy shores along the coast, New Bruns- 
wick to Virginia and inland on sand hills and lake and river shores, west 
to Manitoba and North Dakota. Flowering from May to July. 


Violet Family 
Wiikorllarcieiare 

Because of their abundance and beauty, the violets take rank with the 
favorites among our native flowers. Our species of violets are all low 
herbs, either leafy-stemmed or stemless, that is, the leaves and flowers arising 
directly from the rootstock. The flowers possess five stamens, the two 
lowest with appendages that project into the spur or nectar sac of the lower 
and odd petal. Only these two stamens are developed in the apetalous 
flowers which come after the petaliferous flowers in most stemless species 
except the Bird’s-foot Violet. The petaliferous flowers are somewhat 
irregular, the lower petal extended into a spur or sac back of the flower, the 
lateral petals usually narrower than the two upper petals, and usually only 
the spur and lateral petals adorned at the base in some species with hairs. 

Allied species of some groups freely hybridize when growing together. 
The hybrids commonly display characters more or less intermediate to those 
of the parent species, and show marked vegetative vigor, but very often 
also impaired fertility. These violets of hybrid origin are frequently unlike 
the mother plant and unlike one another, reverting variously in succeeding 
generations to the characters of the two original species. 

Two cultivated species of violets are common in the east, and some- 
times escape from cultivation. They are the English, Marsh or Sweet 
Violet (Viola odorata Linnaeus), and the Pansy or Heartsease 
(Viola tricolor Linnaeus) with variously colored flowers. The 
original form, and the one which the cultivated forms revert to sooner or 


later, has violet or purple flowers. The large Garden Pansy is the product 


[ 


Memoir 15 N. Y. State Museum 


WILD FLOWERS OF NEW YORK 


Plate 132 


A. WOOLLY HUDSONIA; FALSE HEATHER 
Hudsonia tomentosa 


B. DWARF GINSENG OR GROUNDNUT 


Panax trifolium 


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WILD FLOWERS OF NEW YORK Neal 


of various crosses of Viola tricolor with allied species of the 
Old World. In addition to these, there are about thirty-three native 
species of violets in New York, beside the many natural hybrids. The 
following key to the native species may be useful in determining certain 
species not illustrated here. 


I Stemless; the leaves and scapes directly from a rootstock or from runners 
iRetalsibright yellow leaves orbiculam- ee.) eee aeseee eee nm WE wOwuingGls to ili a 
Petals violet, purple or white 
Cleistogamous flowers wanting; petals all beardless; leaves divided............. 
2V. pedata 
Cleistogamous flowers present, at least later in the season 
Rootstock thick, often stout, without stolons; lateral petals bearded 
Cleistogamous flowers ovoid on short prostrate peduncles; their capsules 
mostly purplish 

Leaves except rarely the earliest, palmately five- to eleven-lobed 

or parted; foliage villous-pubescent 
Leaf-lobes blunt, lateral ones broad........ Be Veepraulembanira: 
Leaf-lobes acuminate, lateral ones linear...4 V. perpensa 
Early and late leaves uncut; others three- to seven-lobed or parted 
& Wo wart Iolo 2) 
Leaves all uncut; blades ovate to reniform, cordate, crenate- 

serrate 
Plants nearly or quite glabrous; petals violet-purple; seeds 
brown 
Petioles smooth plantsiofmoist soll eenee eae ee 
6 V. papilionacea 
Petioles glandular roughened; plants of dry soil.......... 
7 V. latiuscula 
Leaves very hairy, especially beneath and on the petioles; 
SOS Chydke [OOM coccanneeodaenecoucuc SW. SOrOria 
Leaves hirsutulous above, otherwise smooth; seeds buff....... 
9 V. hirsutula 
Cleistogamous flowers ovoid on ascending peduncles, soon elongated 

Leaves pubescent beneath and on the petioles; sepals and their 
auricles ciliolate; blades broadly ovate, cordate................. 


to V. septentrionalis 


172 NEW YORK STATE MUSEUM 


Leaves glabrous beneath and on the petioles... ... sie) Wien Ehteaealisa) ss) 
Cleistogamous flowers on erect peduncles, their capsules green 
Leaves broadly ovate, blunt at the apex; sepals obtuse 
Cleistogamous flowers ovoid; spurred petal villous.......... 
12) Va nepih ro pihsyalilea 
Cleistogamous flowers long and slender, spurred petal glabrous 
i We Coie tullAwe 
Leaves lobed or the margins sharply incised or toothed toward the 
subcordate or truncate base; spurred petal villous, lateral ones 
with capillary beard 
Blade of the mature leaves ovate-oblong, ciliate, finely pubes- 
(Goines joes Walks ses 5ngendoands 14 V. fimbriatula 
Blade of the mature leaves lanceolate, usually smooth; petioles 
nein onssla aestent rien nictone aires HS AVeESiaveuht tea biel 
Blade of the mature leaves broadly ovate or deltoid 
Margin coarsely toothed near the base; blades sometimes 
lobed Ree A mice ee eee Ne 16: Vin, ‘emmlan gama ta 
Margin sharply toothed toward the base and more or less 
pectinately incised: 75. -i5----...27) V. pectana ta 
Blade of mature leaves primarily three-lobed or three-parted, 
the segments two to three-cleft into linear or oblanceolate 
lODES Seer ror res eee ee 18 Vi brit tonuania 
Rootstock slender (or thicker and scaly with age); plants usually from 
stolons 
Petals pale violet; leaves minutely hairy on the upper surface; spur 
leneeer oo IhhaS ello. oon aoeM obbs ona eenne LOM Ve, Sel ketaailedis 
Petals white, with dark purple lines on the lower three 
Cleistogamous capsules ovoid, usually purplish; woodland plants 
Leaves reniform, lateral petals beardless; stolons short....... 
20-V. renifolia 
Leaves broadly ovate, acute; lateral petals bearded: seeds 
ObLuseiatathebasest yee eae eet Ve ienicrojomiutiva 
Leaves ovate, acute or acuminate; lateral petals beardless; 
SSNS ACES CIS OSE Ga opangcosandancabe oad We lile@inel A 
Cleistogamous capsules ellipsoid; always green; peduncles erect: 
bog and wet meadow species 


WILD FLOWERS OF NEW YORK 17 


Leaves broadly ovate or orbicular, cordate, obtuse 


23uNe pa lelkemis 

Leaves oblong to ovate, the base slightly cordate to tapering 
24 V. primulifolia 

Leaves lanceolate to elliptical..........25 V. lanceolata 


2 Leafy-stemmed; the flowers axillary 
Style capitate, beakless, bearded near the summit, spur short; stipules nearly entire, 
soon scarious 
Petals yellow 
Sparingly pubescent; root-leaves usually one to three...26 V. eriocarpa 
Markedly pubescent; root-leaves usually wanting... .. ay No jG CSCCms 
Inner face of the petals white with yellow base, outer face usually violet; leaves 
usually broadly ovate, acuminate, subglabrous.........28 V. canadensis 
Style not capitate; spur long; stipules bristly toothed, herbaceous 
Spur 2 to 4 lines long; lateral petals bearded; styles bent at tip, with short beard 
Petals whiteron cream-coloredis... 4.25 aa eee ee OM Veni baler a) 
Petals violet-blue 
Herbage glabrous or nearly so; leaves orbicular or suborbicular 


Stipules ovate-lanceolate, bristly serrate; leaves often 17 inches 


Wit LC. scrcloct = = Wow arc ee 30 V. conspersa 
Stipules linear, entire except at base; leaves not over three-fourths 
Olvanlimchiayicerallpine penn rrene an Via alba diomica 


Herbage puberulent; stems ascending; blades mostly ovate............ 
32 Va adunicia 

Spur 4 to 6 lines long, lateral petals beardless; style straight and smooth........ 
33 V. rostrata 

Style much enlarged upward into a globose, hollow summit; stipules large, leaflike, 
pectinate at base; upper leaves and middle lobe of stipules entire or nearly so; dry 


SAM Gym lACES yeas a ielesilons ar eae 5 SoA eels Eto BAe Ve Gae ielel sic 


Bird’s-foot Violet 


Viola pedata Linnaeus 
Plate 133 
Leaves nearly smooth from a short, stout, erect rootstock, three- 
divided, the lateral divisions pedately three to five-parted or cleft, the 


174 NEW YORK STATE MUSEUM 


segments linear to spatulate, often two to four-cleft or toothed near the 
apex. Early leaves usually smaller and less deeply dissected. Corolla 
two-thirds to 14 inches broad, the upper petals dark violet, the lower three 
lilac-purple, or as in the common northern variety (var. lineariloba 
DeCandolle) here illustrated, all the petals lilac-purple, all beardless, the 
orange tips of the stamens large and conspicuous in the center of the 
flower. Seed pods smooth, green; seeds copper-colored. 

Apetalous or cleistogamous flowers are never formed in this species, 
but petaliferous flowers are frequent in late summer and autumn. 

Common in dry or sandy fields and open woods from Massachusetts 
to Florida and Louisiana, less abundant or locally common inland to 
Minnesota. Flowering in May and June. 


Early Blue or Palmate-leaved Violet 
Viola palmata Linnaeus 
Plate 134a 

Leaves palmately five to eleven-lobed or parted, erect or nearly so, 
from a thick, usually oblique, simple or branched rootstock, the leaf seg- 
ments variously toothed or cleft, the middle segment usually the widest, 
more or less villous beneath, especially on the veins and on the petioles, 
the upper surface of the leaves often smooth; early leaves smaller and 
usually less divided than the later ones. Flowers on stalks about as long 
as the leaves, two-thirds to 1 inch broad, violet-purple; sepals blunt; 
cleistogamous flowers on prostrate peduncles; seeds brown. 

In woods and thickets, rarely in open fields and most abundant in 
rather dry, rich soil on wooded hills, Massachusetts to Minnesota, south 
to Florida. Flowering from April to June. Like several of the other 
blue-flowered violets it is locally known as Johnny-jump-up. 


Among its close relatives in the east is Viola triloba 
Schweinitz, in which the earliest leaves and those put forth in late summer 
are usually with uncut, reniform, cordate blades, the summer leaf blades 
4 to 6 inches wide, densely villous beneath and on the petioles, three-lobed 


or three-parted. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 133 


BIRD’S-FOOT VIOLET 
Viola pedata var. lineariloba 


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WILD FLOWERS OF NEW YORK iS 


Wetroul a perpensa Greene, of the Great Lakes region, possesses 
linear, acuminate lateral leaf lobes. 


Coast or Britton’s Violet 
Viola brittoniana Pollard 
Plate 136b 
Early leaves reniform to ovate in outline, entire or incised, from a 
thick, erect rootstock; mature leaves three-parted and the segments two 
to four-cleft into linear or oblanceolate, acute lobes, the middle lobe some- 
what the widest; smooth except for minute pubescence on the upper surface 
and margin. Flowering scapes as long or longer than the leaves. Flowers 
large, I to 14 inches broad, rich purple with a conspicuous white throat. 
Sepals linear-lanceolate, acuminate. 
In peaty or moist, sandy soil along the coast, southern Maine to Vir- 
ginia. Flowering in May and June. Next to Viola pedata, one of 
the most showy of our native species of violet. 


Woolly Blue Violet; Sister Violet 
Viola sororia Willdenow 
Plate 135a 

Leaves ovate to orbicular from a stout, simple or branched rootstock; 
petioles and under surfaces of the young leaves, and often the scapes, villous- 
pubescent; leaf blades blunt or pointed, heart-shaped, the margins crenate- 
serrate, sometimes becoming 4 inches wide when mature. Corolla violet 
to lavender, and occasionally white; sepals broad, usually blunt, finely 
ciliate below the middle; petals rather broad, the lower and lateral ones 
densely bearded with white toward the center of the flower. Cleistogamous 
flowers ovoid on short, horizontal peduncles, usually underground, but 
lengthening and ascending as the capsule ripens; capsules green, mottled 
with brown; seeds dark brown. 

In rocky or rich woodlands, moist meadows and on shady ledges, 
Quebec to Minnesota, south to North Carolina. Flowering in April 
and May. 


176 NEW YORK STATE MUSEUM 


Quite as abundant is the Meadow or Hooded Blue Violet (Viola 
papilionacea Pursh) with nearly glabrous foliage, very large leaf 
blades which are reniform or ovate, deep-blue flowers, the odd petal often 
narrow and boat-shaped, usually beardless; capsules ellipsoid, green or 
dark purple. Common in moist fields and groves, frequently about 
dwellings. 

The Broad-leaved Wood Violet (Viola latiuscula Greene) 
possesses broadly ovate-deltoid leaf blades, the earliest ones blunt and 
tinged with purple beneath; petioles glandular-roughened; flowers violet- 
purple. In dry, open woods in sand or gravel. 

The Southern Wood Violet (Viola hirsutula Brainerd) enters 
our range only in southern New York. It is a small species with leaves 
lying close to the ground, the blades orbicular to reniform, purplish 
beneath, silvery pubescent above; flowers reddish purple. 

The Northern Wood Violet (Viola septentrionalis Greene) 
is common in moist, open woodlands south to Connecticut and Pennsyl- 
vania. The foliage, except the earliest leaves, is hirsutulous. The leaf 
blades are ovate to reniform, heart-shaped, ciliate and blunt; sepals blunt, 
closely ciliolate nearly to the tip; flowers deep violet to pale lilac. 


LeConte’s Violet 
Viola affinis LeConte 
Plate 135b 

Foliage nearly or quite smooth; rootstocks slender, branching, or the 
plants growing in matted clusters. Early leaves narrowly ovate and heart- 
shaped, more or less long pointed toward the apex, the margins crenate- 
serrate; petioles slender and smooth. Flowers violet with a conspicuous 
white center; the cleistogamous flowers small, on ascending stalks. 

Common in moist meadows, low woodlands and shady borders of 
streams, New England to Wisconsin, south to Georgia and Alabama. 
Flowering from April until June. The mature leaves of midsummer are 
about 2 inches wide. 


The Northern Bog Violet (Viola nephrophylla Greene) 
resembles LeConte’s Violet in some respects, but the leaves are broader 
and more blunt. It occurs in cold, mossy bogs and sometimes along 
borders of streams and lakes from Quebec to British Columbia south to 
Connecticut and Wisconsin. It appears to be abundant in a bog in Bergen 
swamp, Genesee county. 


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Marsh Blue Violet 


Viola cucullata Aiton 
Plate 138b 


Leaves and stems smooth or nearly so; leaf blades, except the earliest, 
broadly ovate to reniform, heart-shaped at the base, the margins strongly 
cucullate (rolled inward) when the leaves are young, the apex pointed, 
margins crenate-serrate; when mature 2 to 4 inches broad. Flowering 
stalks much longer than the leaves. Flowers violet-blue with a dark-blue 
throat or center, or sometimes entirely white; lateral petals bearded, the 
lower or spur petal smooth and usually shorter than the lateral ones. Cleis- 
togamous flowers on long, slender, erect stalks, their capsules green; seeds 
nearly black. 

In moist meadows, springy places in woodlands and along streams, 
Quebec to Georgia. Flowering from late in April until June. 


Ovate-leaved Violet 
Viola fimbriatula J. E. Smith 


Plate 137a 

Rootstock long and stout, sometimes branching. The earliest leaf 
blades ovate and blunt; the later ones oblong-ovate, acute, finely pubescent, 
especially beneath, the margins crenulate toward the apex, the bases usually 
somewhat heart-shaped or truncate and sharply toothed, incised or auricu- 
late. Flowering stalks about as long as the leaves when first in bloom or 
longer than the leaves in later flowers; the corolla violet-purple. Capsules 
green; seeds brown. Cleistogamous flowers on erect peduncles. 

Dry fields and hillsides, throughout the eastern states and south to 
Georgia. Flowering in May and June. 


Arrow-leaved Violet 


Viola sagittata Aiton 


Plate 137b 
Leaves erect from a stout rootstock, smooth, or sometimes ciliate or 


finely pubescent; their petioles longer than the blades which are lanceolate 


178 NEW YORK STATE MUSEUM 


or oblong-lanceolate, 1 to 4 inches long, truncate or heart-shaped at the 
base and blunt or pointed at the apex, the base hastately or sagittately 
toothed or cleft, the earliest leaves often deltoid-ovate, blunt, and merely 
crenate at the base. Flowers on stalks about as long as the leaves, the 
corolla violet-purple. 

Moist banks, fields and wet meadows, Massachusetts to Minnesota, 
south to Georgia and Louisiana. Flowering in May and June. The 
smooth form appears to be the commoner on the coastal plain, while around 
the Great Lakes region and eastward to the Hudson River valley occurs a 


form with pubescent foliage. 


Triangle-leaved Violet 
Viola emarginata (Nuttall) LeConte 
Plate 136a 

Foliage glabrous, succulent, frequently in dense tufts from stout or 
matted rootstocks. Leaf blades at flowering time narrowly ovate or 
triangular, slightly heart-shaped; the later ones broadly ovate or deltoid, 
1 to 3 inches wide, often as broad as long, the base truncate or slightly 
heart-shaped, coarsely toothed or incised toward the base. Flowering 
scapes usually longer than the leaves; flowers violet-blue, the petals often 
notched at the ends. 

Dry woods, hillsides and fields, southern New York southward. Flow- 
ering in April and May. 


Viola emarginata acutiloba Brainerd, found on Staten 
Island, possesses leaf blades (of mature leaves) which are five-cleft or five- 
parted, the middle lobe long and narrow, the lateral ones shorter and 
narrower than the middle lobe. 

Closely related to the Triangle-leaved Violet is the Cut-leaved Violet 
(Viola pectinata Bicknell) in which the blades of the mature 
leaves are ovate-deltoid, wider than long, the margin deeply dentate or 
pectinate with numerous small linear acute, entire lobes. Low meadows 
and edges of salt meadows near the coast. 


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WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum : Plate 138 


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A. MEADOW BEAUTY; DEER GRASS B. MARSH BLUE VIOLET 
Rhexia virginica é Viola cucullata 


WILD FLOWERS OF NEW YORK 179 


Great-spurred or Selkirk’s Violet 
Viola selkirkit Pursh 
Plate 139a 

Leaves and scapes 2 to 4 inches high from a slender rootstock or stolon. 
Leaf blades thin, crenate margined, ovate to suborbicular, deeply heart- 
shaped, the basal lobes converging or overlapping; leaves small at flowering 
time, when mature 1 to 2} inches wide, smooth except for minute, spreading 
hairs on the upper surface of the leaves. Flowers pale violet, not bearded, 
the spur 23 to 4 lines long and much enlarged toward the rounded end. 

In shaded ravines and cool mountain forests, New Brunswick to Penn- 
sylvania and Minnesota, north to Greenland. In central New York its 
favorite habitat is the moss-covered rocks and boulders beneath limestone 
cliffs and shaded by dense forests of mixed hardwoods and hemlock. Flow- 
ering in April and May. 

. Large-leaved White Violet 
Viola incognita Brainerd 
Plate 139b 

Foliage somewhat pubescent with soft, white hairs, especially when 
young, upper leaf surfaces smooth. Leaves ascending from slender root- 
stocks. Leaf blades at flowering time orbicular or reniform, two-thirds to 
2 inches wide, abruptly short pointed at the apex, cordate at the base; 
summer leaves with large, somewhat roughened blades, broadly ovate, 
cordate with an open sinus, acute, 2 to 4 inches wide or larger. Flowers 
white, on stalks as long as the leaves at flowering time, the lateral petals 
bearded, the upper pair obovate; seeds brown. In summer the plants 
produce numerous filiform runners. 

Mountainous and low, moist woodlands, Newfoundland to Dakota and 
south to Tennessee. 


The Kidney-leaved White Violet (Viola renifolia A. Gray) is 
densely pubescent throughout, with reniform leaf blades which are dis- 
tantly crenate-serrate on the margins and rounded at the apex; petals 
white, all beardless, the three lower with brownish veins. In Arbor Vitae 


180 NEW YORK STATE MUSEUM 


swamps and cold woods, Newfoundland to Mackenzie river, south to Penn- 
sylvania and Minnesota and along the Rocky mountains to Colorado. 

The Sweet White Violet (Viola blanda _ Willdenow) has the 
petioles and scapes smooth and often tinged with red; smaller leaves, longer 
flowering stalks with very fragrant white flowers; lateral petals beardless, 
the upper pair of petals often long, narrow and strongly reflexed or some- 
times twisted; seeds dark brown and minutely roughened. 

The Northern White Violet (Viola pallens (Banks) Brainerd) 
has small, broadly ovate or orbicular, smooth, pale-green leaves; scapes 
much longer than the leaves, bearing white, slightly fragrant flowers. 
Common in mossy bogs and wet meadows. 


Primrose-leaved Violet 


Viola primulifolia Linnaeus 


Plate 140a 

Leaf blades oblong to ovate, obscurely crenate-serrate on the margins, 
smooth or somewhat hairy, especially toward the base of the petioles; the 
leaves and flower stalks arising from slender rootstocks or stolons. Flow- 
ering scapes 2 to 10 inches high, usually longer than the leaves. Flowers 
white, the three lower petals purple-veined, the lateral ones shghtly or not 
at all bearded; capsules green; seeds reddish brown. Numerous leafy 
stolons are developed in late summer. 

A frequent violet of moist, open, especially sandy soil near the coast 
from New Brunswick to Florida and Louisiana. Flowering in May 


and June. 
Lance-leaved or Water Violet 


Viola lanceolata Linnaeus 


Plate 140b 

Foliage smooth and plants usually profusely stoloniferous in late 
summer, the stolons rooting at the nodes and bearing numerous apetalous 
flowers; the rootstocks slender. Flowering stalks 2 to 4 inches high or 
higher; mature leaves lanceolate or elliptical in shape, the blade 2 to 6 
inches long and one-eighth to three-fourths of an inch wide, tapering 
gradually below into the margined reddish petiole; margins of the leaves 
obscurely crenulate. Flowers white, the three lower petals striped with 


purplish veins. Fruiting capsules green; seeds dark brown. 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum : Plate 139 


A. GREAT-SPURRED OR SELKIRK’S VIOLET 
Viola selkirkit 


B. LARGE-LEAVED WHITE VIOLET 
Viola incognita 


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WILD FLOWERS OF NEW YORK 181 


Open bogs, marshes and moist meadows, Nova Scotia to Minnesota 
and southward. Flowering in May and June or sometimes as early as the 
latter part of April. 


The Round-leaved Yellow Violet (Viola rotundifolia 
Michaux) possesses oval or orbicular, blunt leaves, heart-shaped with 
repand-crenulate margins; at flowering time about 1 inch wide; in mid- 
summer 2 to 4 inches wide and flat upon the ground; flowers bright yellow, 
the three lower petals with brown lines. In cold woods, Maine to Ontario, 
south to Georgia. Very common in the Adirondack and Catskill mountains. 
In other parts of ‘the State rather rare or local. 


Smoothish Yellow Violet 
Viola eriocarpa Schweinitz 
Plate 134b i 
Commonly with two to four ascending stems from a single rootstock. 
Basal leaves often several, long petioled with ovate to reniform blades, 
smooth except for minute pubescence on the upper part of the stem and 
on the lower leaf surfaces along the veins; the stems bearing one to three 
short-petioled leaves, each broadly ovate, slightly heart-shaped at the 
base and long pointed at the apex, the uppermost ones smaller and nearly 
sessile. Flowers in the upper axils, yellow, the lateral petals bearded. 
Fruiting capsules ovoid, woolly white or rarely nearly smooth; seeds brown. 
In low, open, moist woods, Nova Scotia to Manitoba, south to Georgia 
and Texas. Flowering in May and June. 


The closely related Hairy or Downy Yellow Violet (Viola 
pubescens Aiton) is softly pubescent throughout; stems usually but 
one from a rootstock; usually without basal leaves, but bearing one to three 
leaves on the stem, which are broadly ovate or reniform. In dry, rich 
woods, Nova Scotia to Dakota, south to Virginia and Missouri. 


Canada Violet 
Viola canadensis Linnaeus 
Plate 141 
Stems 6 to 18 inches high, usually several or many together from a 
perennial root, smooth or nearly so. Leaves broadly ovate, heart-shaped, 


182 NEW YORK STATE MUSEUM 


pointed at the apex, the margins toothed. Basal leaves numerous on long 
petioles. Flowers in the axils of the stem leaves, often appearing throughout 
the season from May to July, whitish with a bright-yellow eye, the upper 
petals more or less tinged with violet on the outside, the lower petal striped 
with fine, dark lines. 

In upland and mountainous forests, New Brunswick to Saskatchewan, 
south to South Carolina and Alabama. 


The Pale or Striped Violet (Viola striata Aiton) possesses less 
ascending and more angular stems than V. canadensi5§, 6 to 12 inches 
high when in flower; the flowers white or cream-colored, densely bearded 
in the center. In low and shaded places, New York to Minnesota, south 
to Georgia. 


American Dog Violet 
Viola conspersa Reichenbach 
Plate 142b 

Usually several stems ascending from an oblique, branched rootstock, 
3 to 6 inches long at flowering time. Leaf blades orbicular, heart-shaped, 
crenate-serrate on the margins, blunt, one-half to 2 inches wide, the upper 
ones smaller and more pointed. Flowers numerous, usually pale violet 
or rarely white, raised above the leaves on axillary stalks, 2 to 3 inches 
long. 

Common in low ground or moist, shaded woods, Quebec to Minnesota, 
south to Georgia. Flowering from early spring until late in May. 


Closely related, but more dwarfed, with small, orbicular blades 
and deep violet flowers is the Alpine Dog Violet (Viola labradorica 
Schrank) occurring in New York only on the higher mountains of the 
Adirondacks. 


(The Sand Violet (Viola adunca J..E. smith: Viisubyestura 
Greene) is finely puberulent, the stems only 2 to 6 inches long; leaf 
blades ovate, one-half to 1 inch long, crenulate, blunt, subcordate; flowers 
deep violet, with a straight, blunt spur about 3 lines long. Rather local in 
sandy or sterile soil or on dry stony ridges, Quebec to Maine and westward. 


Wir D ELOW ERS OR INE W. YORK 


Memoir 15 N. Y. State Museum Plate 141 


CANADA VIOLET 
Viola canadensis 


WILD FLOWERS OF NEW YORK 183 


Long-spurred Violet 
Viola rostrata Pursh 


Plate 142a 

Stems usually numerous from an elongated, jointed rootstock, 4 to 8 
inches high. Leaves orbicular to broadly ovate, heart-shaped, nearly or 
quite smooth, serrate on the margins, the upper ones pointed, the lower 
and basal leaves blunt. Petaliferous flowers on long, slender stalks, violet 
with a dark purple-violet center, not bearded, the spur slender and one- 
half of an inch long or longer. 

Shady hillsides and moist woods in leaf mold, Quebec to Michigan, 
south to Georgia. Flowering in May and June. 


The Field Pansy (Viola rafinesquii Greene) is an annual 
plant with slender, smooth, erect stems, 3 to 8 inches high, sometimes 
branched. Leaves small, somewhat rounded on slender petioles; their 
stipules large, conspicuous and deeply cut or fringed. Flowers small, 
bluish white to cream-colored. In fields and open woods, southern New 
York to Michigan and southward to Georgia and Texas. Flowering in 
April and May. 

Loosestrife Family 
Salicariaceae 
(Lythraceae) 

The Swamp Loosestrife and the Spiked or Purple Loosestrife belong 
to this family, which in addition to these two species illustrated here, is 
represented in this State by two smaller flowered species of Lythrum 
(Lythrum hyssopifolia Linnaeus and L. alatum Pursh) and 
the Clammy Cuphea or Blue Waxweed (Parsonsia petiolata 
(Linnaeus) Rusby). 

Swamp Loosestrife; Willow-herb 


Decodon verticillatus (Linnaeus) Elliott 
Plate 143a 


An herblike perennial growing usually in swamps or shallow water. 
Although appearing like an herbaceous plant it is more or less shrubby. 
The stems are angular, recurved, smooth and somewhat woody below, 
3 to 10 feet long, often rooting at the tip when they reach the soil or mud. 
Leaves lanceolate, opposite or verticillate, 2 to 5 inches long, one-third 


184 NEW YORK STATE MUSEUM 


to 1 inch wide, smooth above, somewhat hairy beneath, pointed at both 
ends, on very short petioles. Flowers numerous in cymelike axillary 
clusters; calyx broadly campanulate; corolla about an inch or less broad, 
petals cuneate at the base, pink-purple, the slender filaments of the stamens 
projecting from the flower. Fruiting capsule about one-fourth of an inch 
in diameter or slightly less. 

In swamps, shallow water around the edges of lakes and ponds, or 
along slow streams, often forming thickets, Maine to Florida, west to 
Minnesota, Tennessee and Louisiana. Flowering in June and July. Also 


known as peatweed or slink-weed, wild oleander and grass poly. 


Spiked or Purple Loosestrife 
Lythrum salicaria Linnaeus 
Plate 143b 
Stems four-angled, 2 to 4 feet high or sometimes taller from a perennial 
root, smooth or somewhat pubescent or tomentose above and more or less 
branched. Leaves opposite or sometimes in threes, sessile, lanceolate, 
clasping and heart-shaped at the base 2 to 3 inches long, one-fourth to 
one-half of an inch wide. Flowers purple, one-half to two-thirds of an 
inch long and half as broad, in dense, terminal, branched racemes inter- 
spersed with numerous small leaves; petals four or five, usually five; 
stamens eight or ten, the longer ones scarcely projecting beyond the flower. 
Native of Europe but thoroughly naturalized and common in wet 
places and swamps throughout the east. Very common along the Hudson 
river from Albany to New York. Flowering in July and August. 


Meadow Beauty Family 
Melastomaceae 
Meadow Beauty; Deer Grass 


Rhexia virginica Linnaeus 
Plate 138a 
Stems square, 8 to 18 inches high, the angles of the stem usually slightly 
winged, hairy or nearly smooth; roots perennial and fibrous with a few small 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 142 


4 


A. LONG-SPURRED VIOLET 
Viola rostrata 


B. AMERICAN DOG VIOLET 
Viola conspersa 


nt 


Wi 


WILD FLOWERS OF NEW YORK 


Memoir 15 N. Y. State Museum Plate 143 


l we = 7 


A. SWAMP LOOSESTRIFE; WILLOW-HERB B. SPIKED OR PURPLE LOOSESTRIFE 
Decodon verticillatus Lythrum salicaria 


WILD FLOWERS OF NEW YORK 185 


tubers. Leaves opposite, sessile or nearly so, ascending, ovate or elliptical- 
ovate, pointed at the apex, narrowed or rounded at the base, 1 to 2 inches 
long, one-half to 1 inch wide, with a few scattered hairs on both surfaces, 
conspicuously three to five-nerved, the margins ciliate-serrulate. Flowers 
bright purple, 1 to 14 inches broad, few or several in terminal clusters; 
calyx-tube urn-shaped, constricted above with four triangular-pointed 
lobes, and like the stalk of the flower glandular-pubescent; petals four, 
broadly obovate; stamens eight, equal; anthers yellow, linear, curved and 
minutely spurred on the back. Fruit a four-celled, four-valved capsule 
with numerous small rough, bent seeds. 

In moist, sandy meadows and marshes, Maine to northern New York, 
Ontario and Iowa, south to Florida, Louisiana and Missouri. Flowering 
from July to September. Common on the coastal plain, but rare or local 
inland, except east and north of Oneida lake, where it is very abundant in 
certain places. 


The Maryland Meadow Beauty (Rhexia mariana Linnaeus) 
occurs from Long Island southward. It is more densely hairy, the stems 
are not angled and the leaves are narrower and more spreading. 


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